A checklist of species in the pollen wasp subfamily Masarinae is presented, including synonyms and distributional summaries. The 297 valid species and 15 additional subspecies are arranged in 14 genera, divided into two tribes, Gayellini and Masarini, with the latter divided into the subtribes Priscomasarina, Paragiina, and Masarina. Thirty-two new combinations are made.
INTRODUCTION
The most recent monograph of the Masarinae is by Richards (1962). Since that work there have been numerous changes to the classification of these wasps. The family group arrangement is now different: Richards treated Masarinae in the present sense as a family, with three subfamilies: Euparagiinae, Gayellinae, and Masarinae. Masarinae was divided into tribes, Paragiini and Masarini. Carpenter (1981) reduced Masaridae to a subfamily of Vespidae, removing Euparagiinae, which is the sister-group to all other extant vespids, reducing Richards' Gayellinae and Masarinae to tribes, and sinking Richards' Paragiini and Masarini. Carpenter (1997) reconstituted these latter groups and recognized them as subtribes within Masarini, to which F. W Gess (1998) added another subtribe, Priscomasarina. The generic classification has also undergone considerable change: Giordani Soika (1974), Snelling (1986), and F. W. Gess (1998) each added a genus to the 18 recognized by Richards, while Carpenter (1989), van der Vecht and Carpenter (1990), and Carpenter (1997) synonymized eight. And of course, there have been numerous species descriptions and synonymies published since Richards (1962).
More recently, S. K. Gess (1996) reviewed the Masarinae and provided a list of described species, along with distributional summaries. That list is useful; however, it lacks citations of the original references, limiting its taxonomic value. There have been a number of descriptions and synonymies since that list was published; moreover, preparation of the present checklist has revealed a few errors in both Richards (1962) and S. K. Gess (1996).
The present checklist follows the generic classification in van der Vecht and Carpenter (1990), with one amendment. The generic synonymies in van der Vecht and Carpenter's catalog were based on my ongoing phylogenetic studies in the Vespidae. Only parts of the cladistic study of Masarinae have as yet been published (Carpenter, 1989, 1997; see summary in Carpenter, 1993). Because the rationale for most of the synonymy was not yet published, S. K. Gess (1996) did not follow them in her list. Carpenter (1997) subsequently detailed the basis for generic reclassification in the Paragiina. I will summarize the rationale for the remaining synonymy here.
The monophyly of Trimeria + Microtrimeria is shown by the forewing cu-a distad of the fork of M and CuA by a distance nearly equal to its length, a single occipital carina, the palpal formula 2:3, loss of the lateral pronotal groove, the short pyriform tegula, the posteriorly lamellate scutellum, the female foretarsi with lobes of unique form, the hooked hairs on the female foretarsi, and the compressed mid- and hindcoxae. Both genera are evidently monophyletic, as is the Trimeria subgenus Willinkia Richards, but that does not mean that their recognition is warranted. The characters by which all these taxa differ are for the most part variable within other genera, and so their recognition contributes little to the process of efficient diagnosis (Farris, 1979). Accordingly, they are treated as synonyms.
Richards (1962) recognized two subgenera within Celonites, but whereas Eucelonites is defined by the autapomorphy of an axillary process, the typical subgenus is defined only by its absence. It is evidently paraphyletic, and Eucelonites is thus synonymized with Celonites.
The monophyly of Quartinia + Quartinioides + Quartiniella is shown by the forewing marginal cell appendiculate with a nebulous vein, the forewing cu-a oblique, the hindwing preaxillary excision deep, the single long occipital carina, and the palpal formula 1:3. Both Quartiniella and Quartinioides are primarily based on partitioning a trend in the reduction of wing venation. Quartiniella is defined by the loss of forewing 3rs-m and 2m-cu; Quartinioides has 2m-cu present but attenuate and interrupted, whereas Quartinia has it complete. Other features listed in Richards' (1962: 127) key mostly vary within these taxa or are erroneous (e. g., Quartinioides has two hindtibial spurs). The remainder are consistent with the inference that Quartinia is paraphyletic in terms of Quartinioides, which is in turn paraphyletic in terms of Quartiniella (e. g., Quartinioides is also separated from Quartinia by the preaxillary excision deep, the jugal lobe reduced, and the stipes medially produced, all characters shared with Quartiniella). In any case, the formal taxonomic partitioning of essentially continuous variation should be rejected, and Quartinioides and Quartiniella are accordingly synonymized with Quartinia.
Finally, I previously (van der Vecht and Carpenter, 1990) treated Masarina as a synonym of Jugurtia. Both genera have autapomorphies, but considering the few species of Masarina recognized at that time (i. e., four; see Richards, 1962, Gess and Gess, 1988), I did not regard the features by which the genera differ as significant as those which they share. Further, the recognition of a genus Masarina might be confusing because that name is also used for a subtribe. F. W. Gess (1997) has since described six more species of Masarina and pointed out additional diagnostic features, and I agree that maintaining the genera is now justified. A total of 14 genera (and no subgenera) for the Masarinae is thus recognized here.
The arrangement of the checklist is alphabetical. Each species and subspecies is followed by the original citation and then by synonyms, which are listed chronologically. The status given each taxon generally follows the most recent author to treat that taxon. I do not regard the recognition of subspecies as taxonomically useful, but as such taxa are a mixture of synonyms and valid phylogenetic species, each case should be studied before appropriate corrective action is taken. Misspellings are indicated by exclamation marks in brackets. To facilitate ready reference, each publication is cited in abbreviated form. Where two dates are listed, the first is the true date of publication, so far as can be determined, with the date following in parentheses being the date printed on the publication. The sex of the type specimens, if stated in the description, is then listed. The original locality is quoted for each description, and the type depository is then listed in parentheses. If the sex of the holotype or lectotype is known, this is listed with the depository, except where a single sex was specified in the original description, in which case this information is not repeated, or where syntypes have not had a lectotype designated (a few exceptions to these rules are corrections to original descriptions). A code word is used to designate most type depositories in the checklist, typically the name of city in which the collection is located; this precedes the name of the museum or institution. Museums or institutions where type specimens are kept are as follows.
Berlin: Museum für Naturkunde der Humboldt-Universität, Berlin, Germany
BOL: Biologiezentrum des Oberösterreichischen Landesmuseums Linz, Austria
Bremen: Übersee-Museum, Bremen, Germany
Budapest: Hungarian Natural History Museum, Budapest, Hungary
Buenos Aires: Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” Buenos Aires, Argentina
Cambridge: Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA
Canberra: Australian National Insect Collection, CSIRO, Canberra, Australia
Cape Town: South African Museum, Cape Town, South Africa
Copenhagen: Zoologisk Museum, Kobenhavns Universitet, Copenhagen, Denmark
Davis: Bohart Museum of Entomology, University of California, Davis, California, USA
Geneva: Muséum d'Histoire Naturelle, Geneva, Switzerland
Genova: Museo Civico di Storia Naturale “Giacomo Doria,” Genoa, Italy
Grahamstown: Albany Museum, Grahamstown, South Africa
Hamburg: Zoologisches Museum, Universität Hamburg, Hamburg, Germany
Helsinki: Finnish Museum of Natural History, Helsinki, Finland
IPC: Instituto Pedagógico de Chile [Universidad de Chile, Santiago, Chile]
Leiden: Nationaal Natuurhistorisch Museum, Leiden, the Netherlands
Linz: coll Gusenleitner, Pfitznerstrasse 31, A-4020 Linz/Donau, Austria
London: Natural History Museum, London, England
Madrid: Museo Nacional de Ciencias Naturales, Madrid, Spain
Moscow: Zoological Museum, Moscow State University, Moscow, Russia
New York: American Museum of Natural History, New York, New York, USA
Oxford: Hope Entomological Collections, Oxford University, Oxford, England
Paris: Muséum National d'Histoire Naturelle, Paris, France
Perth: Western Australian Museum, Perth, Australia
Philadelphia: Academy of Natural Sciences, Philadelphia, Pennsylvania, USA
Pretoria: Transvaal Museum, Pretoria, South Africa
St. Petersburg: Zoological Institute, Russian Academy of Sciences, St Petersburg, Russia
San Francisco: California Academy of Sciences, San Francisco, California, USA
San Lorenzo: Museo Nacional de Historia Natural del Paraguay, San Lorenzo, Paraguay
Santiago: Museo Nacional de Historia Natural, Santiago, Chile
Sofia: National Museum of Natural History, Bulgarian Academy of Sciences, Sofia, Bulgaria
Tucumán: Fundación y Instituto Miguel Lillo, Tucumán, Argentina
Turin: Museo di Zoologia Sistematica dell'Università di Torino, Turin, Italy
Uppsala: Zoological Museum, Uppsala University, Uppsala, Sweden
Venice: Museo Civico di Storia Naturale, Venice, Italy
Vienna: Naturhistorisches Museum Wien, Vienna, Austria
Washington: National Museum of Natural History, Smithsonian Institution, Washington, DC, USA
Zürich: Eidgenössiche Technische Hochschule, Zürich, Switzerland
A few smaller private collections are not included in the above list, being mostly collections whose present whereabouts are doubtful.
Distributional data are summarized by country in the final paragraph for each species. An asterisk denotes a new locality record.
SUBFAMILY MASARINAE LATREILLE, 1802
Masarides Latreille, 1802, Hist. Nat. Crust. Insect. 3: 367, family. Type genus: Masaris Fabricius, 1793.
Mazaridinae Schulz, 1911, Zool. Ann. Würzburg 4: 199. Unnecessary emendation of Masarinae Latreille.
Celonitinae Börner, 1919, Biol. Zentralbl. 39(4): 186. Type genus: Celonites Latreille, 1802.
Saryarinae Kuznetzov, 1923, J. Turkestan Branch Russ. Geogr. Soc. 16: 127. Type genus: Saryara Kuznetzov, 1923.
Tribe Gayellini Bradley, 1922
Gayellinae Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 374, 376, 380 (key), subfamily. Type genus: Gayella Spinola, 1851.
Genus Gayella Spinola
Gayella Spinola, 1851, in Gay, Hist. Fis. Polit. Chile, 6: 328, genus. Type species: Gayella eumenoides Spinola, 1851, by monotypy.
Gazella[!]Reed, 1894, Proc. Zool. Soc. London 1893: 685.
araucana Willink
Gayella araucana Willink, 1956, Bol. Acad. Nac. Cienc. 39: 341, 342, 346, figs. 3, 4, female, male –“Chile, Prov. Santiago: Renca” (holotype female, IPC).
Distribution: Chile: Atacama, Coquimbo, Valparaíso, Santiago, O'Higgins, Maule, Biobío.
eumenoides Spinola
Gayella eumenoides Spinola, 1851, in Gay, Hist. Fis. Polit. Chile. Zool. 6: 333, pl. II, fig. 2, male, female –“provincias del norte y sobretodo en Santa Rosa” (lectotype female, Paris).
Gayella sicheliana von Schulthess, 1910, Deutsch. Entomol. Z.: 189, as a synonym of eumenoides, attributed to de Saussure.
Distribution: Chile: Atacama, Coquimbo, Valparaíso, Santiago, O'Higgins, Maule.
luispenai Willink and Ajmat de Toledo
Gayella luispenai Willink and Ajmat de Toledo, 1979, Acta Zool. Lilloana 35: 427, 428, 429, figs. 1, 6, 7, male, female –“Riconada, Jujuy, Argentina” (holotype male, Tucumán).
Distribution: Argentina: Jujuy; Bolivia: Potosí.
mutilloides de Saussure
Gayella mutilloides de Saussure, 1855, Ét. Fam. Vesp. 3: 114, female –“Le Chili” (London).
Gayella odyneroides; von Schulthess, 1910, Deutsch. Entomol. Z.: 189. Lapsus calami.
Gayella mutilloides nigerrima Giordani Soika, 1960 (1958), Boll. Mus. Civ. Stor. Nat. Venezia 11: 80, female –“Cile” (Venice).
Distribution: Chile: Valparaíso, Santiago, Biobío, Araucanía; Argentina: Chubut, Neuquén, Rio Negro.
patagonica Willink
Gayella patagonica Willink, 1956, Bol. Acad. Nac. Cienc. 39: 341, 342, 350, figs. 9–11, female, male –“Esquel, Chubut, Argentina” (holotype female, Tucumán).
Gayella cerceroides Giordani Soika, 1960 (1958), Boll. Mus. Civ. Stor. Nat. Venezia 11: 82, female –“N.W. Patagonia, 1000–3000 piedi” (London).
Distribution: Chile: Santiago; Argentina: Chubut, Mendoza, Rio Negro.
reedi Willink
Gayella reedi Willink, 1963, Acta Zool. Lilloana 19: 385, 1 fig., female –“Casa Blanca, Valparaíso, Chile” (San Francisco).
Distribution: Chile: Atacama, Coquimbo, Valparaíso.
Genus Paramasaris Cameron
Paramasaris Cameron, 1901, Trans. Am. Entomol. Soc. 27: 311, genus. Type species: Paramasaris fuscipennis Cameron, 1901, by monotypy.
Zethoides Cameron, 1904, Trans. Am. Entomol. Soc. 30: 93, genus, non Zethoides Fox, 1899. Type species: Zethoides flavolineatus Cameron, 1904, by monotypy.
Plesiozethus Cameron, 1905, Entomologist 38: 269, replacement name for Zethoides Cameron.
Metazethoides Schulz, 1906, Spolia Hym.: 213, unnecessary replacement name for Zethoides Cameron.
Paragayella Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 87, 89, 99, genus. Type species: Paragayella richardsi Giordani Soika, 1974, by original designation and monotypy.
brasiliensis Giordani Soika
Paramasaris brasiliensis Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 105, female, male –“Brasile: Nova Teutonia, Santa Catarina” (holotype female, Cambridge).
Distribution: Brazil: Santa Catarina; Argentina: Misiones.
cupreus Giordani Soika
Paramasaris cupreus Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 104, 106, female –“Columbia: Caqueta, Florencia, 480 m.” (London).
Distribution: Colombia: Caqueta, Magdalena, Putumayo; Peru: Loreto.
fuscipennis Cameron
Paramasaris fuscipennis Cameron, 1901: 312, female –“Santa Fé Mountains, New Mexico” (London).
Zethoides flavolineatus Cameron, 1904, Trans. Am. Entomol. Soc. 30: 94, male –“Panama (Pacific side)” (London).
Distribution: Colombia; Panama; Guatemala; Mexico: Jalisco, Sinaloa.
richardsi (Giordani Soika)
Paragayella richardsi Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 88, fig. 2, pl. II, 101, female –“Brasile: Mato Grosso, Serra Roncados, R. S. Base Camp” (London).
Distribution: Brazil: Goiás, Mato Grosso.
Tribe Masarini Latreille, 1802
Subtribe Priscomasarina Gess, 1998
Priscomasarina Gess, 1998, J. Hym. Res. 7(2): 296, 299, subtribe. Type genus: Priscomasaris Gess, 1998.
Genus Priscomasaris Gess
Priscomasaris Gess, 1998, J. Hym. Res. 7(2): 296, genus. Type species: Priscomasaris namibiensis Gess, 1998, by original designation and monotypy.
namibiensis Gess
Priscomasaris namibiensis Gess, 1998, J. Hym. Res. 7(2): 296, 300, figs. 2–9, female, male –“Namibia: between Palm and Khorixas (20.17S 14.05 E)” (holotype female, Grahamstown); also from five other localities in Namibia.
Distribution: Namibia.
Subtribe Paragiina Bradley, 1922
Paragiini Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 387 (key), tribe. Type genus: Paragia Shuckard, 1837.
Genus Metaparagia Meade-Waldo
Metaparagia Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 748, genus. Type species: Paragia pictifrons Smith, 1857, by original designation.
Riekia Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 54, genus. Type species: Riekia nocatunga Richards, 1962, by original designation and monotypy.
Rolandia Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 57, genus. Type species: Paragia maculata Meade-Waldo, 1910, by original designation and monotypy.
Ammoparagia Snelling, 1986, Contrib. Sci. 378: 1, 2, genus. Type species: Ammoparagia hua Snelling, 1986, by original designation and monotypy.
angulata (Richards)
Riekia angulata Richards, 1968, J. Aust. Entomol. Soc. 7: 101, female –“QUEENSLAND … Cunnamulla” (Canberra); also from New South Wales.
Distribution: Australia: Queensland, New South Wales.
borreriae (Snelling)
Rolandia borreriae Snelling, 1986, Contrib. Sci. 378: 1, 8–9, 11, figs. 11, 17, 24, male, female –“Cooper Creek (12°06;prS, 133°04;prE), 19 km E by S Mt. Borradaile, Northern Territory” (holotype male, Perth).
Distribution: Australia: Northern Terrritory.
confluens (Snelling)
Paragia confluens Snelling, 1986, Contrib. Sci. 378: 1, 14, fig. 28, female –“Western Australia … 36 km ESE Minnie Creek Homestead, 24°38;prS, 115°42;prE” (Perth).
Distribution: Australia: Western Australia.
doddi Meade-Waldo
Metaparagia doddi Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 748, female –“Cairns, N. Queensland” (London).
Distribution: Australia: Queensland.
houstoni (Snelling)
Rolandia houstoni Snelling, 1986, Contrib. Sci. 378: 1, 8, figs. 9, 12, 14, 15, 18, 22, 23, 26, 29, male, female –“Western Australia … 10 km ESE Meedo Homestead (25°40;prS, 114°37;prW)” (holotype male, Perth).
Distribution: Australia: Western Australia.
hua (Snelling)
Ammoparagia hua Snelling, 1986, Contrib. Sci. 378: 1, 3, 5, figs. 1–8, male, female –“Western Australia … 43 km ENE Landor Homestead (25°08;prS, 116°54;prE)” (holotype male, Perth).
Distribution: Australia: Western Australia.
maculata (Meade-Waldo)
Paragia maculata Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 31, 32, female, male –“Western Australia” (lectotype female, London).
Distribution: Australia: Western Australia.
nocatunga (Richards)
Riekia nocatunga Richards, 1962, Revision. Study Masarid Wasps: 54, figs. 55–59, female –“New South Wales: Nocatunga (7 miles N. of)” (Canberra).
Distribution: Australia: New South Wales.
pictifrons (Smith)
Paragia pictifrons Smith, 1857, Cat. Hym. Brit. Mus.: 2, female –“Swan River” (London).
Distribution: Australia: Western Australia.
Genus Paragia Shuckard
Paragia Shuckard, 1837, Trans. Entomol. Soc. London 2: 81, genus. Type species: Paragia decipiens Shuckard, 1837, by monotypy.
Alpha Dalla Torre, 1904, Genera Insect. 19: 4, name for division I of genus Paragia Shuckard in de Saussure, 1855, Ét. Fam. Vesp. 3: 54 (3 species). Non Alpha de Saussure, 1854.
Beta Dalla Torre, 1904, Genera Insect. 19: 4, name for division II of genus Paragia Shuckard in de Saussure, 1855, Ét. Fam. Vesp. 3: 56 (5 species). Non Beta de Saussure, 1854.
Cygnaea Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 60, subgenus of Paragia Shuckard. Type species: Paragia vespiformis Smith, 1865, by original designation and monotypy.
Paragiella Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 67, subgenus of Paragia Shuckard. Type species: Paragia odyneroides Smith, 1850, by original designation.
australis australis de Saussure
Paragia australis de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xxi, sex not stated –“Nova Hollandia” (Paris).
Distribution: Australia: New South Wales.
australis borealis Richards
Paragia australis borealis Richards, 1962, Revision. Study Masarid Wasps: 67, 71, female, male –“Queensland: 17 miles west of Morven” (holotype female, Canberra).
Distribution: Australia: Queensland.
bicolor de Saussure
Paragia bicolor de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xxi, sex not stated –“Nova Hollandia” (Paris).
Distribution: Australia: New South Wales.
calida Smith
Paragia calida Smith, 1865, Trans. Entomol. Soc. London (3) 2: 392, male –“Adelaide” (London).
Distribution: Australia: South Australia, Western Australia.
deceptor Smith
Paragia deceptor Smith, 1862, Trans. Entomol. Soc. London (3) 1: 56, female –“Australia” (London).
Paragia deceptrix; Schulz, 1906, Spolia Hym.: 213. Unnecessary emendation.
Distribution: Australia: Queensland, New South Wales.
decipiens aliciae Richards
Paragia decipiens aliciae Richards, 1962, Revision. Study Masarid Wasps: 63, 64, female, male –“Northern Territories: Alice Springs” (holotype female, London).
Distribution: Australia: Northern Terrritory, Queensland.
decipiens decipiens Shuckard
Paragia decipiens Shuckard, 1837, Trans. Entomol. Soc. London 2: 82, pl. VIII, fig. 3, female –“New South Wales” (London).
Distribution: Australia: New South Wales, Victoria, South Australia.
generosa Richards
Paragia generosa Richards, 1962, Revision. Study Masarid Wasps: 67, 69, female –“Queensland: 30 miles S.S.W. of Ayr” (Canberra).
Distribution: Australia: Queensland.
hirsuta Meade-Waldo
Paragia hirsuta Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 749, male –“Cairns, N. Queensland” (London).
Distribution: Australia: Queensland.
magdalena Turner
Paragia magdalena Turner, 1908: 89, female –“MACKAY, Queensland” (London).
Distribution: Australia: Queensland.
mimetica Richards
Paragia mimetica Richards, 1968, J. Aust. Entomol. Soc. 7: 101, 102, male, female –“WESTERN AUSTRALIA … 10 m. W. Watheroo” (holotype male, Canberra).
Distribution: Australia: Western Australia.
monocesta Snelling
Paragia monocesta Snelling, 1986, Contrib. Sci. 378: 1, 15–16, figs. 27, 30, 31, female, male –“Western Australia … 10 km ESE Meedo Homestead, 25°40;prS, 114°37;prE” (holotype female, Perth).
Distribution: Australia: Western Australia.
morosa Smith
Paragia morosa Smith, 1868, Trans. Entomol. Soc. London 1868: 251, female –“Champion Bay” (London).
Distribution: Australia: Western Australia.
nasuta Smith
Paragia nasuta Smith, 1868, Trans. Entomol. Soc. London 1868: 252, female, male –“Champion Bay” (lectotype female, London).
Distribution: Australia: Queensland, Western Australia.
odyneroides Smith
Paragia odyneroides Smith, 1850, Trans. Entomol. Soc. London (2) 1: 41, pl. 5, fig. 2, male –“Hunter River, New Holland” (London).
Paragia bidens de Saussure, 1855, Ét. Fam. Vesp. 3: 59, male –“L'Adélaïde, Nouvelle –Hollande” (London).
Paragia praedator de Saussure, 1855, Ét. Fam. Vesp. 3: 59, female –“Le sud de la Nouvelle –Hollande” (London).
Distribution: Australia: Queensland, New South Wales, Victoria, South Australia.
oligomera Snelling
Paragia oligomera Snelling, 1986, Contrib. Sci. 378: 1, 13, female –“Western Australia … 10 km NE Wanneroo” (Perth).
Distribution: Australia: Western Australia.
perkinsi Meade-Waldo
Paragia perkinsi Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 750, female –“Cairns, N. Queensland” (London).
Distribution: Australia: Queensland.
propodealis Richards
Paragia propodealis Richards, 1968, J. Aust. Entomol. Soc. 7: 101, 104, female –“NEW SOUTH WALES … Caldwell” (Canberra).
Distribution: Australia: New South Wales.
schulthessi Turner
Paragia schulthessi Turner, 1936, Ann. Mag. Nat. Hist. (10) 18: 352, female –“Dedari, 40 miles west of Coolgardie, W. Australia” (London).
Distribution: Australia: Western Australia.
smithii de Saussure
Paragia tricolor Smith, 1850, Trans. Entomol. Soc. London (2) 1: 41, male, not female.
Paragia smithii de Saussure, 1854, Ét. Fam. Vesp. 3: pl. II fig. 1, female; 1855, Ét. Fam. Vesp. 3: 55 –“L'Australie Occidentale” (type depository unknown).
Distribution: Australia: New South Wales, Victoria, South Australia, Western Australia.
sobrina Smith
Paragia sobrina Smith, 1869, Trans. Entomol. Soc. London 1869: 309, female –“Champion Bay” (London).
Paragia excellens Smith, 1869, Trans. Entomol. Soc. London 1869: 309, female, male –“Champion Bay” (lectotype female, London).
Distribution: Australia: Western Australia.
tricolor Smith
Paragia tricolor Smith, 1850, Trans. Entomol. Soc. London (2) 1: 41, pl. 5, fig. 1, female not male –“Perth, W. Australia” (London).
Paragia saussurii Smith, 1857, Cat. Hym. Brit. Mus.: 2, female –“Perth” (London).
Paragia saussurei[!]; Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 32; Richards, 1962, Revision. Study Masarid Wasps: 66, 294.
Distribution: Australia: Western Australia.
venusta Smith
Paragia venusta Smith, 1865, Trans. Entomol. Soc. London (3) 2: 393, female –“Swan River” (London).
Paragia concinna Smith, 1868, Trans. Entomol. Soc. London 1868: 251, female –“Champion Bay” (London).
Distribution: Australia: Western Australia.
vespiformis Smith
Paragia vespiformis Smith, 1865, Trans. Entomol. Soc. London (3) 2: 393, female –“Swan River” (London).
Distribution: Australia: Western Australia.
walkeri Meade-Waldo
Paragia walkeri Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 31, 33, male –“Adelaide River, Northern Territory” (London).
Distribution: Australia: Northern Terrritory, Queensland.
Subtribe Masarina Latreille, 1802
Genus Celonites Latreille
Celonites Latreille, 1802, Hist. Nat. Crust. Insect. 3: 368, genus. Type species: Masaris apiformis Fabricius, 1793 (= Vespa abbreviata Villers, 1789), by monotypy.
Chelonites[!] Leach, 1815, “Entomology” in Edinburgh Encyclop. IX(1): 154.
Coelonites[!] Blanchard, 1840, Hist. Nat. Anim. Artic. 3: 387.
Paracelonites Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 108, subgenus of Celonites Latreille (6 species). Unavailable; no type species designated.
Eucelonites Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 112, subgenus of Celonites Latreille (4 species). Unavailable; no Type species: designated.
Eucelonites Richards, 1962, Revision. Study Masarid Wasps: 214, 215, subgenus of Celonites Latreille. Validation by type selection of Eucelonites Kostylev, 1935. Type species: Celonites cyprius de Saussure, 1854, by original designation.
abbreviatus abbreviatus (Villers)
Vespa abbreviata Villers, 1789, C. Linn. Entomol. 3: 281, sex not stated –“Europa. In Gallia Australiori” (destroyed).
Chrysis dubia Rossi, 1790, Fauna Etrusca 2: 77, pl. VII, figs. x–xi, sex not stated [female] –“Provinciis Florentina et Pisana” (destroyed?).
Cimbex vespiformis Olivier, 1790, Encyc. Méth.: 772, sex not stated –“en France, en Italie … aux environs de Paris” (destroyed?).
Masaris apiformis Fabricius, 1793, Entomol. Syst. 2: 284, sex not stated –“in Italia” (Copenhagen).
Celonites abbreviatus var. Hungaricus Mocsáry, 1877, Termesz. Füzetek. 1: 90, female –“Hungaria centrali” (Budapest?).
Celonites abbreviatus var. engadinensis von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 139, male, female –“Schweiz. Alpengebiet, Ober-Engadin, Sils Baselgia” (Zürich).
Distribution: Germany; Austria; Hungary; Switzerland; France; Spain; Portugal; Italy; Croatia; Albania; Bulgaria; Greece: Corfu; Cyprus; Turkey; Israel; Morocco.
abbreviatus tauricus Kostylev
Celonites abbreviatus tauricus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 108, female (in subgenus Celonites) – Crimea] (St. Petersburg?).
Distribution: Ukraine.
afer Lepeletier
Celonites afer Lepeletier, 1841, Hist. Nat. Insect. Hym. 2: 585, female –“Oran” (Paris?).
Distribution: Morocco; Algeria; Tunisia; Libya.
andrei Brauns
Celonites Andréi Brauns, 1905, Ann. Mus. Natl. Hung. 3: 228, female –“Terra Capensis: Willowmore” (Pretoria).
Distribution: South Africa: Western Cape, Eastern Cape.
bergenwahliae Gess
Celonites bergenwahliae Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, figs. 8–9, 12–13, 91, female, male –“Cape Province: Clanwilliam District, Klein Alexandershoek (32°20;pr20;dpS, 18°46;prE)” (holotype female, Grahamstown); also from 5 km W of Clanwilliam.
Distribution: South Africa: Western Cape.
capensis Brauns
Celonites capensis Brauns, 1905, Ann. Mus. Natl. Hung. 3: 231, female –“Willowmore” (Pretoria).
Distribution: South Africa: Western Cape, Eastern Cape.
clarus Gusenleitner
Celonites clarus Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, 66, female –“Iran, Teheran, Steppa presso Chitgar, 18 km W Teheran” (Venice).
Distribution: Iran.
clypeatus Brauns
Celonites clypeatus Brauns, 1913, Entomol. Mitt., Berlin 2: 206, female –“Willowmore” (Pretoria).
Distribution: Namibia*; South Africa: Northern Cape*, Eastern Cape.
crenulatus Morawitz
Celonites crenulatus Morawitz, 1888, Horae Soc. Entomol. Ross. 22: 267, female –“in territorio transcaspico (Kiltitschinar)” (St. Petersburg?).
Distribution: Iran*; Turkmenistan.
cyprius cyprius de Saussure
Celonites cyprius de Saussure, 1854, Mém. Soc. Phys. Hist. Nat. Genève 14: 4, male –“l'Ille de Chypre” (Geneva?).
Distribution: Cyprus.
cyprius smyrnensis Richards
Celonites cyprius smyrnensis Richards, 1962, Revision. Study Masarid Wasps: 218 (key), 232, female, male –“Turkey: near Smyrna, Burnova” (holotype female, London); also from Israel (Greece: Rhodes mentioned in key).
Distribution: Greece: Thessalia, Pelopponisos, Rhodes; Turkey; Armenia; Iran; Syria; Lebanon; Israel.
davidi Gess
Celonites davidi Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, fig. 14, 92, male –“Cape Province: Namaqualand, Anenous (29°15′30″S, 17°34′45″E)” (Grahamstown).
Distribution: South Africa: Northern Cape.
discretus Gusenleitner
Celonites discretus Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, figs. 3a, 4b, 62, female –“Iran, Fars, Monti steppiti 5 km N di Persepolis” (Venice).
Distribution: Iran.
fischeri Spinola
Celonites fischeri Spinola, 1838, Ann. Soc. Entomol. France 7: 505, female –“Egypte” (type depository unknown).
?Celonites savignyi de Saussure, 1855, Ét. Fam. Vesp. 3: 91 –“Description de l'Egypte. Hymen. Pl. IX, fig. 19, ♀♂” (type depository unknown).
Distribution: Algeria; Tunisia; Libya; Egypt; Cyprus; Israel; Saudi Arabia.
foveolatus foveolatus Kostylev
Celonites foveolatus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 111, 112, male, female (in subgenus Paracelonites) – [Zakaspian desert] (Moscow, St. Petersburg?).
Distribution: Turkmenistan.
foveolatus nigrior Richards
Celonites foveolatus nigrior Richards, 1962, Revision. Study Masarid Wasps: 218 (key), 234, female –“Israel: Jericho” (Oxford).
Distribution: Turkey; Israel.
gariepensis Gess
Celonites gariepensis Gess, 1997, J. Hym. Res. 6: 36, 41, figs. 1–6, female, male –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from two other localities in Richtersveld.
Distribution: South Africa: Northern Cape.
guichardi Richards
Celonites guichardi Richards, 1962, Revision. Study Masarid Wasps: 218 (key), 233, female –“Libya: Fezzan, El Manakh (50 m. N.W. of Edri)” (London).
Distribution: Libya.
hamanni Gusenleitner
Celonites hamanni Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, 65, 66, fig. 6, female, male –“Türkei Mut” (holotype female, Linz).
Celonites hammani[!]; Yildirim and Özbek, 1997, Atatürk. Ü. Zir. Fak. Der. 28 (2): 224.
Distribution: Turkey; Syria; Jordan.
hellenicus Gusenleitner
Celonites hellenicus Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 109, 110 (key), 112, male, female (in subgenus Eucelonites) –“Hellas: Ano Karmes, (Likeo Bgl.)” (holotype male, BOL); also from elsewhere in Greece.
Distribution: Greece.
humeralis Richards
Celonites humeralis Richards, 1962, Revision. Study Masarid Wasps: 219, 236, female –“Cape Province: near Calvinia, Bloukrans” (London); also from Nieuwoudtville.
Distribution: South Africa: Northern Cape.
hystrix Kostylev
Celonites hystrix Kostylev, 1940, Bull. Soc. Nat. Moscou (Biol.) 49: 138, female, male (in subgenus Eucelonites) –“Kondara, Chaîne Ghissar, 30 km vers le N de Stalinabad (autrefois Duchambé), 1 100 m au –d. du niv. de la mer” (coll. Gussakovskii, St. Petersburg).
Distribution: Tajikistan.
jousseaumei jousseaumei du Buysson
Celonites jousseaumei du Buysson, 1906, Rev. Entomol., Caen 25: 104, female, male –“Obock, Djibouti, Tadjourah” (Paris).
Celonites Joussaumei[!]; von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 140.
Celonites jousseaumei asrensis Giordani Soika, 1957, Brit. Mus. Exped. S.-W. Arabia 1(31): 472, female –“YEMEN: ‘Asr, about 3 miles west of San'a, 8100 ft.” (London).
Distribution: Algeria; Sudan; Dijibouti; Israel; Saudi Arabia; Yemen; Oman; United Arab Emirates; Qatar.
jousseaumei senegalensis Richards
Celonites jousseaumei senegalensis Richards, 1962, Revision. Study Masarid Wasps: 216 (key), 224, female, male –“Senegal: Bambey” (holotype female, Paris).
Distribution: Senegal; Mali; Djibouti.
kostylevi Panfilov
Celonites kostylevi Panfilov, 1961, Arch. Mus. Zool. Univ. Moscou 8: 165, female, male (in subgenus Celonites) – [Kirghizia: east bank of Issyk-Kul’ lake, forest valley Ak-Ulen] (Moscow).
Distribution: Kyrgyzstan.
kozlovi Kostylev
Celonites kozlovi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 110 (in subgenus Paracelonites) – [Gobi Desert, Mongolia] (St. Petersburg?).
Distribution: Mongolia.
laetus Panfilov
Celonites laetus Panfilov, 1968, Arch. Zool. Mus. Moscow State Univ. 11: 40, fig. 3, female – [Transcaucasia: Nakhichevan’, vicinity of Dzhulfa, desert foothills] (Moscow).
Distribution: Azerbaijan.
latitarsis Gess
Celonites latitarsis Gess, 1992, J. Hym. Res. 1: 141, figs. 1–5, female, male –“Cape Province: Clanwilliam District, 11 km W of Clanwilliam on road to Graafwater” (holotype female, Grahamstown).
Distribution: South Africa: Western Cape.
lobeliae Gess
Celonites lobeliae Gess, 1997, J. Hym. Res. 6: 36, 46, figs. 13–14, female, male –“Cape Province: Betw[een] Nieuwoudtville and top of Vanrhyns Pass” (holotype female, Grahamstown).
Distribution: South Africa: Northern Cape.
longipilis Gusenleitner
Celonites longipilis Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 59, figs. 3b, 4a, female –“Iran, Fers [Fars], Monti steppici 5 km N di Persepolis” (Venice).
Distribution: Iran.
mayeti Richards
Celonites mayeti Richards, 1962, Revision. Study Masarid Wasps: 222 (key), 243, figs. 202, 208, 211, female, male –“France: Bouches-du-Rhône, Camargue” (holotype male, Paris); also from other localities in France and Spain.
Distribution: France; Spain.
michaelseni von Schulthess
Celonites Michaelseni von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land.-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 137, male –“Windhuk” (Hamburg?).
Distribution: Namibia.
modestus biinterruptus Kostylev
Celonites modestus biinterruptus Kostylev, 1940, Bull. Soc. Nat. Moscou (Biol.) 49: 138, female, male (in subgenus Paracelonites) –“Ruidacht, 40 km. de Stalinabad” (coll. Gussakovskii, St. Petersburg).
Celonites modestus bisinterruptus[!]; Richards, 1962, Revision. Study Masarid Wasps: 247.
Distribution: Tajikistan.
modestus modestus Kostylev
Celonites modestus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 110 (in subgenus Paracelonites) – [Pamir, Mountains of Peter the Great] (St. Petersburg?).
Distribution: Tajikistan.
montanus Mocsáry
Celonites montanus Mocsáry, 1906, Ann. Hist.-Nat. Mus. Hung. 4: 198, male –“Turkestan: Alai montes” (Budapest).
Distribution: Kyrgyzstan; China: Xinjiang.
nursei Dover
Celonites nursei Dover, 1925 (1924), J. Proc. Asiat. Soc. Bengal (n. ser.) 20(6): 289, fig. 1, female –“Quetta” (London).
Distribution: Pakistan.
octoannulatus hissaricus Kostylev
Celonites octoannulatus hissaricus Kostylev, 1940, Bull. Soc. Nat. Moscou (Biol.) 49: 137, female, male (in subgenus Paracelonites) –“Ruidacht, 40 km vers le N de Stalinabad (autrefois Duchambé). 3000 m. au-dessus du niv. de la m.” (coll. Gussakovskii, St. Petersburg).
Distribution: Tajikistan.
octoannulatus octoannulatus (Kuznetzov)
Saryara octoannulata Kuznetzov, 1923, J. Turkestan Branch Russ. Geogr. Soc. 16: 127, female, male –“Montes Ac-Tasch, distr. Taschkent, Turkestan. Alt 7500” (St. Petersburg).
Distribution: Uzbekistan.
osseus Morawitz
Celonites osseus Morawitz, 1888, Horae Soc. Entomol. Ross. 22: 268, female –“in territorio transcaspico (Tschikischljar)” (St. Petersburg?).
Distribution: Armenia; Turkmenistan.
peliostomi Gess
Celonites peliostomi Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, figs. 1–5, female, male –“Cape Province: Namaqualand, Springbok, Hester Malan Nature Reserve” (holotype female, Grahamstown).
Distribution: South Africa: Northern Cape.
persicus Richards
Celonites persicus Richards, 1962, Revision. Study Masarid Wasps: 221 (key), 241, female –“South West Persia: K. Sefid” (London).
Distribution: Iran.
phlomis Gusenleitner
Celonites phleomis[!]; Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, fig. 3c. Incorrect original spelling.
Celonites phlomis Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 61, 63, fig. figs. 4c, 5, female, male –“Türkei, Urfa” (holotype female, Linz).
Distribution: Turkey.
pictus pictus Richards
Celonites pictus Richards, 1962, Revision. Study Masarid Wasps: 217 (key), 226, figs. 176–177, male –“Senegal: Bambey” (Paris).
Distribution: Senegal; Morocco; Saudi Arabia; Oman.
pictus rufiventris Gusenleitner
Celonites pictus rufiventris Gusenleitner, 1992, Linz. Biol. Beitr. 24(1): 59, 60, female –“Iran, Bandar Abbas (Strand)” (Linz).
Distribution: Iran.
promontorii Brauns
Celonites promontorii Brauns, 1905, Ann. Mus. Natl. Hung. 3: 232, female –“Willowmore” (Pretoria).
Distribution: South Africa: Northern Cape, Eastern Cape, Free State.
purcelli Brauns
Celonites Purcelli Brauns, 1905, Ann. Mus. Natl. Hung. 3: 226, female, male –“Terra Capensis: Willowmore” (Pretoria).
Celonites purcelli var. flavomaculatus Brauns, 1913, Entomol. Mitt., Berlin 2: 205, male –“Willowmore” (Pretoria).
Distribution: South Africa: Eastern Cape.
rothschildi du Buysson
Celonites rothschildi du Buysson, 1906, Rev. Entomol., Caen 25: 105, male –“Afrique orientale anglaise: Lesammise, Rendilé” (Paris).
Celonites Rotschildi[!]; von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land.-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 140.
Distribution: Kenya; Iran?
rudesculptus Kostylev
Celonites rudesculptus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 109 (in subgenus Paracelonites) – [Armenia] (St. Petersburg?).
Distribution: Armenia.
rugiceps Bischoff
Celonites rugiceps Bischoff, 1928, Abhand. Naturwiss. Verein. Bremen 27 (1): 86, female –“Kreta” (Berlin); also from mainland Greece; Turkey.
Distribution: Croatia; Yugoslavia; Bulgaria; Greece: Argos, Crete; Cyprus; Turkey.
semenovi Kostylev
Celonites semenovi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 112 (in subgenus Eucelonites) – [Persia] (St. Petersburg?).
Distribution: Iran.
spinosus Gusenleitner
Celonites spinosus Gusenleitner, 1966, Polskie Pismo Entomol. 36: 359, fig. 2, male, female –“Kusadasi” (holotype female, Linz); also from other localities in Turkey.
Celonites abbreviatus invitus Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 58, fig. 2b, female, male –“Türkei, Gürün” (holotype female, Linz); also from elsewhere in Turkey, and Armenia.
Distribution: Turkey; Armenia.
tristiculus karatauicus Kostylev
Celonites tristiculus karatauicus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 111, male (in subgenus Paracelonites) –[Kara Tau] (St. Petersburg?).
Distribution: Kazakhstan.
tristiculus tristiculus Kostylev
Celonites tristiculus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 111, female, male (in subgenus Paracelonites) – [Fergana. Syr Darya] (Moscow, St. Petersburg?).
Distribution: Uzbekistan.
tumidiscutellatus Gess
Celonites tumidiscutellatus Gess, 1997, J. Hym. Res. 6: 36, 44, figs. 7–12, female, male –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from three other localities in Namaqualand, and Eastern Cape.
Distribution: South Africa: Northern Cape, Eastern Cape.
turneri Richards
Celonites turneri Richards, 1962, Revision. Study Masarid Wasps: 221 (key), 238, figs. 195–197, male –“Cape Province: Aliwal North” (London).
Distribution: South Africa: Western Cape,* Eastern Cape.
varipennis Richards
Celonites varipennis Richards, 1962, Revision. Study Masarid Wasps: 219 (key), 239, fig. 200, female, male –“Libya: Tibesti, Enneri Meche” (holotype male, London).
Distribution: Libya.
wahlenbergiae Gess
Celonites wahlenbergiae Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, figs. 6–7, 10–11, 87, female, male –“Cape Province: Clanwilliam District, Clanwilliam Dam” (holotype female, Grahamstown).
Distribution: South Africa: Western Cape.
wheeleri Brauns
Celonites Wheeleri Brauns, 1905, Ann. Mus. Natl. Hung. 3: 230, female, male –“Terra Capensis: Willowmore” (Pretoria).
Celonites wheeleri var. immaculatus Brauns, 1905, Ann. Mus. Natl. Hung. 3: 230, female –“Terra Capensis: Willowmore” (Pretoria).
Distribution: Namibia*; South Africa: Eastern Cape.
yemenensis ethiopicus Richards
Celonites yemenensis ethiopicus Richards, 1962, Revision. Study Masarid Wasps: 217 (key), 228, female –“Ethiopia: Diredawa” (London).
Distribution: Ethiopia.
yemenensis yemenensis Giordani Soika
Celonites yemenensis Giordani Soika, 1957, Brit. Mus. Exped. S.-W. Arabia 1(31): 472, fig. 1, male, female –“YEMEN: Ta'izz, on road to Mocha, 4100 ft.” (holotype male, London).
Celonites yemenensis yemensis[!]; Richards, 1969, J. Nat. Hist. 3: 82.
Distribution: Saudi Arabia; Yemen.
zavattarii Giordani Soika
Celonites zavattarii Giordani Soika, 1944, Atti Inst. Ven. Sci. Lett. Art. 103(2): 149, female –“Etiopia: Asile” (lectotype female, Venice); also from Caschei.
Distribution: Ethiopia.
Genus Ceramiopsis Zavattari
Ceramiopsis Zavattari, 1910, Ann. Mus. Civ. Stor. Nat. Genova 44: 531, genus. Type species: Ceramiopsis gestroi Zavattari, 1910, by monotypy.
gestroi Zavattari
Ceramiopsis gestroi Zavattari, 1910, Ann. Mus. Civ. Stor. Nat. Genova 44: 533, figs. A, B, female –“Urucú, Corumbà (Matto Grosso)” (Genova).
Distribution: Bolivia: Santa Cruz; Brazil: Mato Grosso do Sul; Paraguay: Guairá, Paraguarí, San Pedro; Argentina: Formosa, Santa Fé, Entre Rios.
paraguayensis Bertoni
Ceramiopsis paraguayensis Bertoni, 1921, Rev. Soc. Cient. Paraguay 1: 11, female, male –“Asunción, ‘Quinta Caballero’” (San Lorenzo).
Distribution: Bolivia: Santa Cruz; Paraguay: Alto Paraguay, Presidente Hayes, Central, Paraguarí; Argentina: Formosa.
Genus Ceramius Latreille
Ceramius Latreille, 1810, Consid. Gén. Crust. Arach. Insect.: 329, 438. Type species: Ceramius fonscolombei Latreille, 1810, by original designation.
Gnatho Klug, 1810, Mag. Ges. Naturf. Fr. Berl. 4: 32, 36, genus. Non Illiger, 1807 (Coleoptera). Type species: Gnatho lichtensteinii Klug, 1810, by monotypy.
Ceramias[!] Westwood, 1840, Introd. Modern Class. Insect. 2: 237.
Ceramus[!] Westwood, 1840, Introd. Modern Class. Insect. 2: 243.
Paraceramius de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III; 1855, Ét. Fam. Vesp. 3: 67, division of genus Ceramius Latreille (4 species). Type species: Ceramius spiricornis de Saussure, 1854, by subsequent designation of Ashmead, 1902, Can. Entomol. 34: 220.
Ceramioides de Saussure, 1854, Ét. Fam. Vesp. 3: pl. IV no. 1; 1855, Ét. Fam. Vesp. 3: 71, division of genus Ceramius Latreille; 1855: 71, division III of genus Ceramius. Type species: Ceramius cerceriformis de Saussure, 1854, by monotypy.
Euceramius Dalla Torre, 1904, Genera Insect. 19: 5, name for division I of Ceramius Latreille in de Saussure, 1854, Ét. Fam. Vesp. 3: pl. II, 1855, Ét. Fam. Vesp. 3: 62 (3 species). Type species: Ceramius fonscolombei Latreille, by subsequent designation of van der Vecht and Carpenter, 1990, Zool. Verh., Leiden 260.
auctus (Fabricius)
Liris aucta Fabricius, 1804, Syst. Piezat.: 229, sex not stated –“in Tangier” (female Copenhagen).
Ceramius spiricornis de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xx. Nomen nudum.
Ceramius spiricornis de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III, fig. 5, male; 1855, Ét. Fam. Vesp. 3: 68 –“L'Europe méridionale, la France, l'Espagne” (Paris).
Distribution: Morocco; Algeria.
beaumonti (Giordani Soika)
Paraceramius beaumonti Giordani Soika, 1957, Bull. Soc. Sci. Nat. Phys. Maroc 37: 168, figs. 1, 3, 5, male, female –“Asni” (holotype male, Venice); also from elsewhere in Morocco.
Distribution: Algeria; Morocco.
beyeri Brauns
Ceramius Beyeri Brauns, 1903, Z. Syst. Hym. Dipt. 3(2): 69, female, male –“Willowmore … Steytlerville in der Karroo des Caplandes” (Pretoria).
Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.
bicolor (Thunberg)
Philanthus bicolor Thunberg, 1815, Nova Acta Reg. Soc. Sci. Uppsala 7: 127, 131, 289, sex not stated (male, Uppsala).
Ceramius Karrooensis Brauns, 1902, Z. Syst. Hym. Dipt. 2(5–6): 282, female, male –“Willowmore im Capland” (Pretoria).
Ceramius karooensis[!]; Brauns, 1910, Z. Wiss. Insektenbiol. 6(11): 387.
Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.
bischoffi Richards
Ceramius bischoffi Richards, 1963, Zool. Meded. 38: 216, fig. 2, male, female –“Spain: Tornavacas, prov. Cáceres, 1200 m” (holotype male, Leiden); also from elsewhere in Spain.
Distribution: Spain.
braunsi Turner
Ceramius braunsi Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 291 (key), 294, fig. 2, male, female –“Olifant's River, between Citrusdal and Clanwilliam” (holotype male, Cape Town).
Distribution: South Africa: Western Cape, Eastern Cape.
brevitarsis Gess
Ceramius brevitarsis Gess, 1997, J. Hym. Res. 6: 36, 37, female, male –“Cape Province: Namaqualand, Richtersveld N[ational] Park, bet[ween] hills (28.10S, 17.02E), NW Koeroegabvlakte” (holotype female, Grahamstown).
Distribution: South Africa: Northern Cape.
bureschi Atanassov
Ceramius bureschi Atanassov, 1938, Mitt. K. Naturwiss. Inst. Sofia 11: 114, figs. 1, 2a, 3a, 4a, 5a, 6a, male –“Bulgarien, im Park des Palais Wrana bei Sofia” (Sofia).
Ceramius bureschi lycaonius Blüthgen, 1952, Mitt. Münch. Entomol. Ges. 42: 18, male, female –“Attika” (holotype male, Berlin).
Distribution: Bulgaria; Greece; Turkey.
caffer de Saussure
Ceramius caffer de Saussure, 1855, Ét. Fam. Vesp. 3: 76, female –“Cap de Bonne-Espérance” (Torino).
Ceramius consobrinus de Saussure, 1855, Ét. Fam. Vesp. 3: 77, female –“Cap de Bonne-Espérance” (Torino).
Distribution: South Africa: Western Cape.
capensis de Saussure
Ceramius capensis de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III, figs. 3–3a, female; 1855, Ét. Fam. Vesp. 3: 66 –“Cap de Bonne-Espérance” (Turin?).
Distribution: South Africa: Western Cape.
capicola Brauns
Ceramius capicola Brauns, 1902, Z. Syst. Hym. Dipt. 2(5): 278, female, male –“Willowmore” [“Hier”] (Pretoria).
Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape, Free State.
caucasicus André
Ceramius Caucasicus André, 1884, Spec. Hym. Europe Algérie II: 820, male –“Caucase” (Paris?).
Distribution: Iran; Armenia; Turkey.
cerceriformis de Saussure
Ceramius cerceriformis de Saussure, 1853, Ann. Soc. Entomol. France (3), Bull. II: xxi, male –“Cap de Bonne-Espérance” (Torino).
Ceramius vespiformis de Saussure, 1855, Ét. Fam. Vesp. 3: 79, female –“Cap de Bonne-Espérance” (Torino).
Ceramius Schulthessi Brauns, 1902, Z. Syst. Hym. Dipt. 2 (3): 182, female –“Willowmore, Capland” (Pretoria).
Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.
clypeatus Richards
Ceramius clypeatus Richards, 1962, Revision. Study Masarid Wasps: 83, 88 (key), 99, male –“Clan William” (Pretoria).
Distribution: South Africa: Western Cape.
damarinus Turner
Ceramius damarinus Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 292 (key), 293, male, female –“Ongandjera, S.W. Africa … Kamanyab” (holotype male, Cape Town).
Distribution: Namibia.
fonscolombei fonscolombei Latreille
Ceramius fonscolombei Latreille, 1810, Consid. Gén. Crust. Arach. Insect.: 438, sex not stated [male] –“aux environs d'Aix” [p. 329] (Paris).
Distribution: France; Spain; Portugal.
fonscolombei oraniensis Lepeletier
Ceramius Oraniensis Lepeletier, 1841, Hist. Nat. Insect. Hym. 2: 591, female, male –“Oran” (Paris).
Ceramius Doursii André, 1884, Spec. Hym. Europe Algérie II: 818, female, male –“Algérie” (type depository unknown).
Distribution: Morocco; Algeria.
gessi Mauss
Ceramius maroccanus gessi Mauss, 1998, Beitr. Hym.-Tag. Stuttgart (2–4.10.1998): 18. Nomen nudum.
Ceramius gessi Mauss, 1999, Entomol. Scand. 30: 323, 331 (key), 340, figs. 1c, 2c, 3c, 3f, i, 4c, 4g, 4j, 5d–f, 7b, 8c, 9c, 10–11, female, male –“Morocco, 10 km W Tiznit, 29°43;prN 09°50;prW” (holotype male, BOL); also from elsewhere in Morocco.
Distribution: Morocco.
hispanicus Dusmet
Ceramius hispanicus Dusmet, 1909, Act. Mem. Primer Congr. Nat. Españ., Zaragoza, 1908: 179, 180, female, male –“Madrid! y Montarco! (en Madrid). –Jaramiel! (en Valladolid)” (Madrid).
Distribution: Spain.
jacoti Richards
Ceramius nigripennis de Saussure; Brauns, 1913, Entomol. Mitt., Berlin 2: 201, pl. 2, fig. 3, male. Misidentification.
Ceramius jacoti Richards, 1962, Revision. Study Masarid Wasps: 83, 89 (key), 101, male, female –“Cape Province: Ladismith” (holotype male, London); also from Montagu.
Distribution: South Africa: Western Cape.
lichtensteinii (Klug)
Gnatho Lichtensteinii Klug, 1810, Mag. Ges. Naturf. Fr. Berlin 4: 38, pl. 1 figs. 3, e, f, female –“Cap” (Berlin?).
Ceramius macrocephalus de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III figs. 2, 2a, 2c, female; 1855, Ét. Fam. Vesp. 3: 76 –“Cap de Bonne-Espérance” (Paris).
Ceramius rufomaculatus Cameron, 1906, Trans. South African Philos. Soc. 16(4): 325, female –“Pearston, Cape Colony” (London).
Distribution: Namibia?; South Africa: Northern Cape, Western Cape, Eastern Cape, Free State.
linearis Klug
Ceramius linearis Klug, 1824, Entomol. Monogr.: 227, male –“Prom. bonae spei, Distr. Vitenhagen” (Berlin).
Ceramius fumipennis Brauns, 1902, Z. Syst. Hym. Dipt. 2(5): 275, female, male –“Karroosteppe und … Port Elizabeth nahe der Küste” (Pretoria).
Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.
lusitanicus Klug
Ceramius lusitanicus Klug, 1824, Entomol. Monogr.: 223, 230, sex not stated –“Lusitania” (Berlin).
Ceramius lusitanicus var. luteo-clypeata Dusmet, 1909, Act. Mem. Primer Congr. Nat. Españ., Zaragoza, 1908: 179, 180, male –“Montarco! (en Madrid)” (Madrid).
Distribution: Spain; Portugal; Gibraltar.
maroccanus (Giordani Soika)
Paraceramius maroccanus Giordani Soika, 1957, Bull. Soc. Sci. Nat. Phys. Maroc 37: 170, figs. 6, 8, 11–12, female, male –“Mogador” (holotype male, Venice); also from elsewhere in Morocco.
Distribution: Morocco.
metanotalis Richards
Ceramius metanotalis Richards, 1962, Revision. Study Masarid Wasps: 83, 86 (key), 106, fig. 114, female –“Cape Province: Calvinia” (London); also from Nieuwoudtville.
Distribution: South Africa: Northern Cape, Western Cape.
micheneri Gess
Ceramius micheneri Gess, 1968, Novos Taxa Entomol. 57: 3, figs. 1–6, male, female –“Cape Province: Citrusdal” (holotype male, Cape Town); also from Olifants River between Klawer and Clanwilliam.
Distribution: South Africa: Western Cape.
montanus Gusenleitner
Ceramius montanus Gusenleitner, 1990, Linz. Biol. Beitr. 22(2): 565, 567, fig. 2, male, female –“Marokko, Hoher Atlas, Oukaimeden, 2600–2800 m” (Linz); also from elsewhere in Morocco.
Distribution: Morocco.
nigripennis de Saussure
Ceramius nigripennis de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III figs. 4, 4c, female; 1855, Ét. Fam. Vesp. 3: 69 –“Cap de Bonne-Espérance” (Torino).
Ceramius hessei Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 291 (key), 296, fig. 1, male, female –“Kamieskroon, Namaqualand” (holotype male, Cape Town).
Distribution: South Africa: Northern Cape, Western Cape.
palaestinensis (Giordani Soika)
Paraceramius palaestinensis Giordani Soika, 1957, Bull. Soc. Sci. Nat. Phys. Maroc 37: 172, female –“Palestine: Ein Geb” (Venice).
Ceramius palestinensis[!]; Yildirim and Özbek, 1997, Atatürk. Ü. Zir. Fak. Der. 28 (2): 225; Yildirim and Kojima, 1999, Nat. Hist. Bull. Ibaraki Univ. 3: 20.
Distribution: Israel; Syria; Turkey.
peringueyi Brauns
Ceramius Péringueyi Brauns, 1913, Entomol. Mitt., Berlin 2: 194, female –“in der Nähe der Capstadt” (Pretoria).
Distribution: South Africa: Western Cape.
rex de Saussure
Ceramius rex de Saussure, 1855, Ét. Fam. Vesp. 3: 75, female –“Cap de Bonne-Espérance” (Turin?).
Distribution: South Africa: Western Cape, Northern Cape.
richardsi Gess
Ceramius peringueyi Brauns; Richards, 1962, Revision. Stud. Masarid. Wasps: 100, male. Misidentification.
Ceramius richardsi Gess, 1965, Ann. S. Afr. Mus. 48: 219, 221, male, female –“Cape Province: Clanwilliam” (holotype male, Pretoria); also from Paleisheuwel.
Distribution: South Africa: Western Cape.
rubripes Gusenleitner
Ceramius maroccanus rubripes Gusenleitner, 1990, Linz. Biol. Beitr. 22(2): 565, 566, female –“Marokko, Anezal, Col de Tizin Bachoom, 1500 m, 7°3W 30°7N” (coll. H. Tussac); also from elsewhere in Morocco.
Distribution: Morocco.
socius Turner
Ceramius socius Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 291 (key), 297, male, female –“Worcester, Cape Province” (holotype male, London); also from elsewhere in Western Cape.
Distribution: South Africa: Western Cape.
toriger von Schulthess
Ceramius toriger von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 383, female –“South Africa, Cape Colony … St. Rhynspass” (London); also from elsewhere in Northern Cape.
Distribution: South Africa: Northern Cape.
tuberculifer de Saussure
Ceramius tuberculifer de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xx. Nomen nudum.
Ceramius tuberculifer de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III fig. 6, male; 1855, Ét. Fam. Vesp. 3: 69, as a synonym of lusitanicus Klug –“L'Espagne, le midi de la France” (Paris).
Ceramius lusitanicus Klug; de Saussure, 1855, Ét. Fam. Vesp. 3: 69, male, female (in part). Misidentification.
Distribution: France; Spain.
vechti Richards
Ceramius vechti Richards, 1963, Zool. Meded. 38: 213, fig. 1, male, female –“Spain: Tornavacas, prov. Cáceres, 1200 m” (holotype male, Leiden); also from elsewhere in Spain.
Distribution: Spain.
Genus Jugurtia de Saussure
Jugurtia de Saussure, July 1854, Ét. Fam. Vesp. 3: 19, 37, 42–44, generic and specific characters of J. numidica and J. oraniensis; pl. I fig. 4, antenna of J. oraniensis; genus. Type species: Celonites oraniensis Lepeletier, 1841, by subsequent designation of Ashmead, 1902, Can. Entomol. 34: 220.
Jugurthia[!] André, 1886, Spec. Hym. Europe Algérie II, Cat. Méthod. Syn.: 47.
Jugurthia Dalla Torre, 1894, Cat. Hym. 9: 5; E. Saunders, 1905, Trans. Entomol. Soc. London: 399. Unnecessary emendation of Jugurtia de Saussure, 1854.
Jujurtha[!] Ashmead, 1902, Can. Entomol. 34: 220.
Masariella Brauns, 1905, Ann. Mus. Natl. Hung. 3: 223, genus. Type species: Masariella alfkeni du Buysson, 1904, by monotypy.
Ceramiellus von Schulthess, 1922, Deutsch. Entomol. Z. 1922: 401, genus. Type species: Ceramiellus braunsi von Schulthess, 1922, by monotypy.
Juguartia[!] Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 68.
alfkeni (du Buysson)
Masaris alfkeni du Buysson, 1904, Bull. Soc. Entomol. France: 144, female –“Damara land, Swakopmund” (Bremen).
Distribution: Namibia.
algerica (von Schulthess)
Masariella? algerica von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 499 (key), 501, fig. 2, female –“Algeria occidentalis, Ain Sefra, … Ain Fezza” (Zürich).
Distribution: Morocco; Algeria; Libya.
biskrensis Bequaert
Jugurtia biskrensis Bequaert, 1937, Bull. Ann. Soc. Entomol. Belg. 68: 343, 345 (key), 358, fig. 5A–F, female, male –“Algérie. –Biskra” (holotype female, Cambridge).
Distribution: Morocco; Algeria.
braunsi (von Schulthess)
Ceramiellus braunsi von Schulthess, 1922, Deutsch. Entomol. Z. 1922: 402, 2 figs., female, male –“Südwestafrika: Groß- und Klein-Namaland, Grootfontein … Steinkopf” (Berlin).
Distribution: Namibia; South Africa: Western Cape, Eastern Cape.
braunsiella (von Schulthess)
Masaris braunsiella von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 326, fig. 1, male –“Cape Province, Matjesfontein” (London).
Distribution: Namibia; South Africa: Northern Cape, Western Cape, Eastern Cape.
calcarata Richards
Jugurtia calcarata Richards, 1962, Revision. Study Masarid Wasps: 251 (key), 260, male, female –“S. Africa, Cape Province: Worcester” (holotype male, London); also from elsewhere in Western Cape.
Distribution: South Africa: Western Cape.
codoni Gess
Jugurtia codoni Gess, 1997, J. Hym. Res. 6: 36, 53, female, male –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from three other localities in Richtersveld.
Distribution: South Africa: Northern Cape.
confusa Richards
Jugurtia confusa Richards, 1962, Revision. Study Masarid Wasps: 249, 251 (key), 258, figs. 213–215, 218, 220–221, female, male –“S. Africa: Cape Province, Willowmore” (holotype male, London).
Distribution: South Africa: Eastern Cape.
discrepans (Brauns)
Masaris discrepans Brauns, 1913, Entomol. Mitt., Berlin 2: 203, pl. II fig. 9, female, male –“Worcester im Süden der Kapkolonie … und bei Oudtshoorn (Kapland)” (Pretoria).
Distribution: Namibia; South Africa: Western Cape, Eastern Cape.
dispar (Dufour)
Celonites dispar Dufour, 1851, Ann. Soc. Entomol. France (2) 9: 58, 59, pl. 3 figs. 1–8, female, male –“environs de Madrid” (Paris?).
Distribution: Spain; Portugal; Gibraltar.
duplicata Richards
Jugurtia duplicata Richards, 1962, Revision. Study Masarid Wasps: 249, 254 (key), 263, male –“Cape Province: Van Rhynsdorp” (Pretoria).
Distribution: South Africa: Western Cape.
eburnea (Turner)
Masariella turneri eburnea Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 299, fig. 3, female, male –“Kamieskroon, Namaqualand” (holotype male, Cape Town).
Jugurtia polita Richards, 1962, Revision. Study Masarid Wasps: 251 (key), 256, female, male –“S. Africa, Cape Province: Calvinia” (holotype male, London); also from elsewhere in Cape Province.
Distribution: South Africa: Northern Cape.
escalerae Meade-Waldo
Jugurtia escalerae Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 33, female –“S.W. Persia” (London).
Distribution: Turkey; Iran.
eurycara Kostylev
Jugurtia eurycara Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 94 (key), 97, figs. 6–7, male, female –“Caucasus, Armen.” (St. Petersburg); also from Erivanj [Yerevan].
Distribution: Turkey; Iran; Armenia.
irana Kostylev
Jugurtia irana Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 94 (key), 99, female, male –“Via Karvandar-Bampur, S.-O. Persien … Tshanshen-Sadk, Bampur, S.-O. Perasien … Kuusha-Larumba, Bampur, S.-O. Persien … Kurljut-Langer, Chorasan, O. Persien” (St. Petersburg).
Distribution: Iran.
jemenensis Kostylev
Jugurtia jemenensis Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 94 (key), 99, female –“Jemen, Sanaa” (Moscow).
Jugurtia hoggarica Giordani Soika, 1954, Boll. Mus. Civ. Stor. Nat. Venezia 7: 20, male –“Hoggar, Atakor, Guelta Afilale” (Venice).
Jugurtia jemenesis[!]; Richards, 1962, Revision. Study Masarid Wasps: 265.
Distribution: Algeria; Israel; Saudi Arabia; Yemen; Oman; United Arab Emirates.
koeroegabensis Gess
Jugurtia koeroegabensis Gess, 1997, J. Hym. Res. 6: 36, 55, female, male –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from one other locality in Richtersveld.
Distribution: South Africa: Northern Cape.
nadigorum Bequaert
Jugurtia nadigorum Bequaert, 1937, Bull. Ann. Soc. Entomol. Belg. 68: 343, 344 (key), 347, fig. 2A–F, female, male –“Maroc. –Marrakech” (holotype female, Cambridge); also from one other locality.
Distribution: Morocco.
oraniensis (Lepeletier)
Celonites Oraniensis Lepeletier, 1841, Hist. Nat. Insect. Hym. 2: 586, female, male –“Oran” (Paris?).
Jugurtia numidica de Saussure, 1854, Ét. Fam. Vesp. 3: pl. V fig. 3, male; 1855, Ét. Fam. Vesp. 3: 85 –“L'Algérie” (Paris).
Jugurtia numida[!]; de Saussure, 1855, Ét. Fam. Vesp. 3: 85.
Distribution: Morocco; Algeria; Tunisia.
saussurei (Brauns)
Masaris Saussurei Brauns, 1905, Ann. Mus. Natl. Hung. 3: 219, female, male –“Terra Capensis: Willowmore” (Pretoria).
Distribution: South Africa: Western Cape, Eastern Cape.
simpsoni Meade-Waldo
Jugurtia simpsoni Meade Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 448, female –“Patchari … N'jau” (London).
Masariella? testaceopicta von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 499 (key), 502, fig. 3, female –“N. Nigeria, Zungeru” (London).
Distribution: Gambia; Senegal; Burkina Faso; Nigeria.
spinolae (de Saussure)
Masaris spinolae de Saussure, 1855, Ét. Fam. Vesp. 3: 95, female –“Cap de Bonne-Espérance” (Torino).
Jugurtia spinolai[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 92.
Jugurtia spirolai[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 100.
Distribution: South Africa: Western Cape.
tigrina Gess
Jugurtia tigrina Gess, 1997, J. Hym. Res. 6: 36, 50, figs. 1–6, female, male –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from five other localities in Northern Cape.
Distribution: South Africa: Northern Cape.
turneri (von Schulthess)
Masariella? turneri von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 499 (key), 500, fig. 1, female –“Cape Province, Little Carroo, 38 m. E. of Ceres” (London).
Distribution: South Africa: Western Cape.
zarudnyi Kostylev
Jugurtia zarudnyi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 93 (key), 94, figs. 4–5, male, female –“Chorasan … Kermän, Prov. Mokran” (St. Petersburg).
Distribution: Iran.
Genus Masarina Richards
Masarina Richards, 1962, Revision. Study Masarid Wasps: vii, 213, 268, genus. Type species: Masarina familiaris Richards, 1962, by original designation.
ceres Gess
Masarina ceres Gess, 1997, J. Hym. Res. 6: 36, 60, male, figs. 23–24 –“Cape Province: 17 km N of Ceres, near top of Gydo Pass” (Grahamstown).
Distribution: South Africa: Western Cape.
familiaris Richards
Masariella spinolae (de Saussure); Bequaert, 1928, Ann. Mag. Nat. Hist. (10) 2: 187. Misidentification.
Masarina familiaris Richards, 1962, Revision. Study Masarid Wasps: 268, 270, figs. 225–231, male, female –“Cape Province: Camps Bay” (holotype male, London); also from elsewhere in Cape Province.
Masarina communis; Richards, 1962, Revision. Study Masarid Wasps: 269. Lapsus calami.
Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.
hyalinipennis Richards
Masarina hyalinipennis Richards, 1962, Revision. Study Masarid Wasps: 268, male –“Cape Province: van Rhynsdorp” (Pretoria).
Distribution: South Africa: Northern Cape, Western Cape.
mixta Richards
Masarina mixta Richards, 1962, Revision. Study Masarid Wasps: 268 (key), 271, female –“S. Africa, Cape Province: Nieuwoudtville” (London).
Distribution: South Africa: Northern Cape, Western Cape.
mixtoides Gess
Masarina mixtoides Gess, 1997, J. Hym. Res. 6: 36, 61, female, male, fig. 32 –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from two other localities in Richtersveld.
Distribution: South Africa: Northern Cape.
namaqua Gess
Masarina namaqua Gess, 1997, J. Hym. Res. 6: 36, 65, female, male, figs. 25–27, 33 –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from three other localities in Namaqualand.
Distribution: South Africa: Northern Cape.
parvula Gess
Masarina parvula Gess, 1997, J. Hym. Res. 6: 36, 65, female, male, figs. 28–30, 34 –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from two other localities in Namaqualand.
Distribution: South Africa: Northern Cape.
peliostomi Gess
Masarina peliostomi Gess, 1997, J. Hym. Res. 6: 36, 67, female, male, figs. 35–39 –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown).
Distribution: South Africa: Northern Cape.
strucki Gess
Masarina strucki Gess, 1988, Ann. Cape Prov. Mus. (Nat. Hist.) 16: 351, 352, female –“[Cape Province:] Namaqualand, [Springbok]” (Grahamstown).
Distribution: South Africa: Northern Cape; Western Cape.
tylecodoni Gess
Masarina tylecodoni Gess, 1997, J. Hym. Res. 6: 36, 69, female, male, figs. 40–43 –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown).
Distribution: South Africa: Northern Cape.
Genus Masaris Fabricius
Masaris Fabricius, 1793, Entomol. Syst. II: vi, 283, 284, genus (2 species). Type species: Masaris vespiformis Fabricius, 1793, by subsequent monotypy; Latreille, 1802, Hist. Nat. Crust. Insect. 3: 368, placed the second species in a new genus, Celonites.
Massaris Billberg, 1820, Enum. Insect.: 109. Unnecessary emendation of Masaris Fabricius.
Mazaris Schulz, 1911, Zool. Ann. Würzburg 4: 199. Unnecessary emendation of Masaris Fabricius.
Saryara Kuznetzov, 1923, J. Turkestan Branch Russ. Geograph. Soc. 16: 127, genus (2 species). Type species: Saryara longicornis Kuznetzov, 1923, by subsequent designation of van der Vecht and Carpenter, 1990, Zool. Verh., Leiden 260.
aegyptiacus aegyptiacus Meade-Waldo
Masaris vespiformis aegyptiacus Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 447, fig. 2, male –“Cairo, Egypt” (London).
Distribution: Egypt; Israel; Jordan.
aegyptiacus arabicus Giordani Soika
Masaris vespiformis arabicus Giordani Soika, 1957, Brit. Mus. Exped. S.-W. Arabia 1(31): 472, female –“SAUDI ARABIA: Jebel Shammar Province, Hâil district” (London).
Distribution: Saudi Arabia.
carli von Schulthess
Masaris saussurei Carl, 1921, Rev. Suisse Zool. 28: 41, male, female –“Zarawchan; … Sutkend. Turkestan” (Geneva). Non Brauns, 1905.
Masaris carli von Schulthess, 1922, Deutsch. Entomol. Z. 1922: 404, replacement name for saussurei Carl.
Masaris smirnovi Kostylev, 1925, Entomol. Mitt. 14: 150 (key), 152, fig. 2, male –“Perovsk (Turkestan, Syr-Darjja-Gebiet)” (Moscow).
Distribution: Iran; Kazahkstan; Uzbekistan.
gussakovskii Kostylev
Masaris gussakovskii Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 87, 89, fig. 2, female –“Turkestan, distr. Chodzhent” (coll. Gussakovskii, St. Petersburg); also from Konstantinovsk.
Distribution: Uzbekistan.
longicornis (Kuznetzov)
Saryara longicornis Kuznetzov, 1923, J. Turkestan Branch Russ. Geogr. Soc. 16: 127, male –“Montes Ac-Tasch. Alt. 8000′” (St. Petersburg).
Distribution: Afghanistan; Uzbekistan; Kyrgyzstan.
tianshanicus Panfilov
Masaris tianshanicus Panfilov, 1968, Arch. Zool. Mus. Moscow State Univ. 11: 38, fig. 2, female – [Tian Shan: south bank of Issyk-Kul’ lake, Smaller Dzhargylchak (near Akterek), dry hills with steppe vegetation] (Moscow).
Distribution: Kyrgyzstan.
vespiformis Fabricius
Masaris vespiformis Fabricius, 1793, Entomol. Syst. 2: 283, sex not stated –“in Barbariae floribus” (Copenhagen).
Erynnis romandi de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xix, male –“Barbaria et Ægypta” (Paris).
Ceramius hylaeiformis Dalla Torre, 1894, Cat. Hym. 9: 4, as a synonym of vespiformis, attributed to Klug.
Distribution: Morocco; Algeria; Egypt.
Genus Pseudomasaris Ashmead
Pseudomasaris Ashmead, 1902, Can. Entomol. 34: 221, genus. Type species: Masaris occidentalis Cresson, 1871, by original designation.
Toryna Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 407, 411, subgenus of Pseudomasaris Ashmead. Type species: Masaris vespoides Cresson, 1863, by original designation and monotypy.
Holopticus Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 408, 414, subgenus of Pseudomasaris Ashmead. Type species: Masaris texanus Cresson, 1871, by original designation.
Cotyledon Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 408, 430, subgenus of Pseudomasaris Ashmead. Type species: Masaris edwardsii Cresson, 1872, by original designation.
basirufus Rohwer
Pseudomasaris zonalis basirufus Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 452, female –“Death Valley, California” (Washington).
Pseudomasaris bariscipus Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 409 (key), 418, figs. 77, 85, male –“ARIZONA: Quartzite” (Philadelphia).
Pseudomasaris bariscapus[!]; Bequaert, 1940, Bull. Brooklyn Entomol. Soc. 35: 38.
Distribution: USA: California, Nevada, Arizona.
cazieri Bohart
Pseudomasaris cazieri Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 289 (key), 290, male –“Palos Colorados, Durango, Mexico, 8,000 feet elevation” (New York).
Distribution: Mexico: Durango, Chihuahua.
coquilletti Rohwer
Pseudomasaris coquilletti Rohwer, 1911, Proc. U.S. Natl. Mus. 40: 555, female, male –“Los Angeles County, California” (holotype female, Washington).
Distribution: USA: Oregon, California, Utah, Arizona.
edwardsii (Cresson)
Masaris edwardsii Cresson, 1872, Trans. Am. Entomol. Soc. 4: 87, female, male –“California” (Philadelphia).
Distribution: USA: Washington, Oregon, Idaho, Wyoming, Colorado, Utah, Nevada, California, Arizona; Mexico: Baja California Norte.
macneilli Bohart
Pseudomasaris macneilli Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 285 (key), 291, male, female –“Near Sonora Peak (above Sonora Pass), Tuolumne County, California, 11,000 feet elevation” (holotype male, San Francisco); also from elsewhere in California, and Utah.
Distribution: USA: California, Utah.
macswaini Bohart
Pseudomasaris macswaini Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 286 (key), 301, male, female –“Andreas Canyon, Riverside County, California” (holotype male, Davis); also from other localities in California.
Distribution: USA: California; Mexico*: Baja California Norte.
maculifrons (Fox)
Masaris maculifrons Fox, 1894, Proc. Calif. Acad. Sci. (2) 4: 12, female –“El Paraiso, L. Cal.” (San Francisco).
Pseudomasaris albifrons Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 451, male –“New Mexico … Las Cruces” (Washington).
Pseudomasaris zonalis neomexicanus Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 452, female –“Aztec, New Mexico” (Washington).
Pseudomasaris rohweri Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 409 (key), 417, male (in subgenus Holopticus) –“ARIZONA: Quartzite” (Philadelphia); also from Phoenix.
Distribution: USA: California, Nevada, Utah, Arizona, New Mexico; Mexico: Sonora, Baja California Norte.
marginalis (Cresson)
Masaris marginalis Cresson, 1864, Proc. Entomol. Soc. Philadelphia 3: 677, female –“Rocky Mountains, Colorado Territory” (Philadelphia).
Distribution: Canada: British Columbia, Alberta.; USA: Washington, Idaho, Wyoming, Colorado, Nevada, Utah, California, Arizona, New Mexico.
micheneri Bohart
Pseudomasaris micheneri Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 284, figs. 1, 14, 21, 286–287 (key), 298, male, female –“Westgard Pass, Inyo County, California, 6,000 feet elevation” (holotype male, Davis); also from elsewhere in California.
Distribution: USA: California, Nevada.
occidentalis (Cresson)
Masaris occidentalis Cresson, 1871, Trans. Am. Entomol. Soc. 3: 348, female –“Texas” (Philadelphia).
Distribution: USA: Kansas, Texas, New Mexico.
phaceliae Rohwer
Pseudomasaris phaceliae Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 450, male, female –“New Mexico, Albuquerque” (holotype male, Washington); also from Mesilla and Filmore Canyon.
Distribution: USA: Texas, New Mexico, Arizona.
texanus (Cresson)
Masaris texanus Cresson, 1871, Trans. Am. Entomol. Soc. 3: 348, male, female –“Texas” (Philadelphia).
Distribution: USA: Texas.
vespoides (Cresson)
Masaris vespoides Cresson, 1863, Proc. Entomol. Soc. Philadelphia 2: 69, pl. 4, male, female –“Pike's Peak” (Philadelphia).
Pseudomasaris vespoides robertsoni Cockerell, 1913, Proc. Entomol. Soc. Wash. 15: 107, female –“Redlands, California” (Washington).
Distribution: USA: Nebraska, South Dakota, Montana, Washington, Oregon, Idaho, Wyoming, Colorado, Utah, Nevada, California, Arizona, New Mexico; Mexico: Baja California Norte.
wheeleri Bequaert
Pseudomasaris wheeleri Bequaert, 1929, Psyche 36: 61, fig. 1, male, female –“CALIFORNIA … Palm Springs, Riverside Co.” (holotype male, Cambridge).
Distribution: USA: California; Mexico: Baja California Norte.
zonalis (Cresson)
Masaris zonalis Cresson, 1864, Proc. Entomol. Soc. Philadelphia 3: 674, female, male –“Rocky Mountains, Colorado Territory” (Philadelphia).
Pseudomasaris zonalis albopictus Bohart, 1950, Proc. Biol. Soc. Wash. 63: 78, male, female –“Jenny Lake, Grand Teton National Park, Wyoming” (holotype male, San Francisco); also from five other localities in Idaho, Oregon and Washington.
Distribution: Canada: British Columbia; USA: Nebraska, Montana, Washington, Oregon, Idaho, Wyoming, Colorado, Utah, Nevada, California.
Genus Quartinia André
Quartinia “Gribodo, in litteris” André, 1884, Spec. Hym. Europe Algérie II: 811, 822, genus. Type species: Quartinia dilecta André, by monotypy.
Juartinia[!] Cameron, 1904, Z. Syst. Hym. Dipt. 4: 89.
Quartiniella von Schulthess, 1929, Ann. Mag. Nat. Hist. 10(3): 504, 510, genus. Type species: Quartiniella waterstoni von Schulthess, 1929, by monotypy.
Quartinioides Richards, 1962, Revision. Study Masarid Wasps: 127, 170, genus. Type species: Quartinia capensis Kohl, 1898, by original designation.
affinis Richards
Quartinia affinis Richards, 1962, Revision. Study Masarid Wasps: 131 (key), 143, female –“Algeria: Sidi Farouch” (London).
Distribution: Algeria.
albopicta (Richards), new combination
Quartinoides[!] albopicta Richards, 1982, Boll. Mus. Civ. Stor. Nat. Venezia 32: 199, female –“S.W. Africa: Gobabeb, 25°36;prS, 15°10;prE” (Copenhagen).
Distribution: Namibia.
alcestis Richards
Quartinia alcestis Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 164, female, male –“Cape Province: Worcester” (holotype female, London).
Distribution: South Africa: Western Cape.
andromeda (Richards), new combination
Quartinioides andromeda Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 182, female –“Cape Province: Prince Albert Road” (London); also from Matjesfontein.
Distribution: South Africa: Western Cape.
antennata von Schulthess
Quartinia antennata von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 385 (key), 388, fig., male –“South Africa, Cape Colony, Worcester” (London).
Distribution: South Africa: Western Cape.
antigone (Richards), new combination
Quartinioides antigone Richards, 1962, Revision. Study Masarid Wasps: 174 (key), 190, figs. 157–161, female, male –“Cape Province: Matjesfontein” (holotype male, London); also from elsewhere in Western Cape.
Distribution: South Africa: Western Cape.
araxana Giordani Soika
Quartinia araxana Giordani Soika, 1960, Boll. Soc. Entomol. Ital. 90: 133, female, male –“Caucaso: Araxesthal” (lectotype female, Venice).
Distribution: Armenia; Turkey; Iran.
arsinoe (Richards), new combination
Quartinioides arsinoe Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 182, female –“S.W. Africa: Aus” (London); also from South Africa.
Distribution: Namibia; South Africa: Western Cape.
artemis Richards
Quartinia artemis Richards, 1962, Revision. Study Masarid Wasps: 128 (key), 156, female –“Cape Province: Calvinia” (London).
Distribution: South Africa: Northern Cape.
atra von Schulthess
Quartinia atra von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 505, female –“Cape Province, Montagu” (London).
Distribution: South Africa: Western Cape.
atrata Gusenleitner
Quartinia atrata Gusenleitner, 1998, Linz. Biol. Beitr. 30(2): 503, 511, female –“Iran, 15 km SE Sarvestan/Fars, 1800 m” (Linz).
Distribution: Iran.
basuto (Richards), new combination
Quartinioides basuto Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 184, female, male –“Basutoland: Mamathes” (holotype male, London).
Distribution: Lesotho.
breyeri Richards
Quartinia breyeri Richards, 1962, Revision. Study Masarid Wasps: 130 (key), 157, female, male –“Cape Province: N.E. Zoutp. district” [actually Northern Transvaal] (holotype female, Pretoria).
Distribution: South Africa: Northern Transvaal.
canariensis Blüthgen
Quartinia canariensis Blüthgen, 1958, Comment. Biol. 18(6): 6, 11, female –“Fuerteventura: Matural” (Helsinki).
Distribution: Spain: Canary Islands: Fuerteventura.
candida Gusenleitner
Quartinia candida Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 137, 147, female –“Israel: 45 km N Elat, Sand tunes [dunes] E Qetura, 29°58′/35°06′” (coll. Schmid-Egger).
Distribution: Israel.
capensis Kohl
Quartinia capensis Kohl, 1898, Termesz. Füzetek 21: 365, female, pl. XV fig. 14 –“Capland … Algoa-Bai” (Vienna).
Quartinia scutellimacula von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 508, male, female –“Cape Province, Mossel Bay” (lectotype female, London); also from elsewhere in Western Cape.
Distribution: South Africa: Western Cape, Eastern Cape.
ceres (Richards), new combination
Quartinioides ceres Richards, 1962, Revision. Study Masarid Wasps: 177 (key), 206, female, male –“Cape Province: Little Karroo, 38 m. E. of Ceres” (holotype male, London); also from elsewhere in Cape Province.
Distribution: South Africa: Western Cape, Eastern Cape.
chlorotica (Morawitz)
Jugurtia chlorotica Morawitz, 1888, Horae Soc. Entomol. Ross. 22: 270, female –“in territorio transcaspico (Tschikischljar)” (St. Petersburg?).
Distribution: Turkmenistan.
cincta Benoist
Quartinia cincta Benoist, 1929, Bull. Soc. Sci. Nat. Maroc 9: 93, fig. 2, female, male –“Maroc: Asni, dans la vallée de la Reraya, … (altitude, 1.200 m.)” (Paris).
Distribution: Morocco.
cressida (Richards), new combination
Quartinioides cressida Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 203, female, male –“Cape Province: Camps Bay, Cape Peninsula” (holotype male, London); also from elsewhere in Cape Province, and Namibia.
Distribution: Namibia; South Africa: Northern Cape, Western Cape, Eastern Cape.
cretica Gusenleitner
Quartinia cretica Gusenleitner, 1994, Linz. Biol. Beitr. 26(1): 331, female –“Griechenland: Kreta, Kastro W Ammoudari; 35.18N 24.10E; 1000–1200 m” (Linz); also from elsewhere in Crete.
Distribution: Greece: Crete.
cyllene (Richards), new combination
Quartinioides cyllene Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 201, female, male –“Cape Province: 13 miles N.E. of Touws R.” (holotype female, London); also from Worcester.
Distribution: South Africa: Western Cape.
cynara (Richards), new combination
Quartinioides cynara Richards, 1962, Revision. Study Masarid Wasps: 173, 174 (key), 189, female –“Cape Province: Willowmore” (Pretoria); also from Prince Albert Road.
Distribution: South Africa: Western Cape, Eastern Cape.
diana (Richards), new combination
Quartinioides diana Richards, 1962, Revision. Study Masarid Wasps: 171 (key), 178, female –“S.W. Africa: Aus” (London).
Distribution: Namibia.
dilecta André
Quartinia Dilecta “Gribodo (in litteris)” André, 1884, Spec. Hym. Europe Algérie II: 824, pl. XLV figs. 3, 7, female, male –“Tunis” (type depository unknown).
Distribution: Morocco; Algeria; Tunisia.
dryope (Richards), new combination
Quartinioides dryope Richards, 1962, Revision. Study Masarid Wasps: 173, 175 (key), 196, male –“Cape Province: Willowmore” (London).
Distribution: South Africa: Eastern Cape.
elissa (Richards), new combination
Quartinioides elissa Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 204, female, male –“Cape Province … Worcester” (holotype male, London); also from elsewhere in Western Cape.
Distribution: South Africa: Western Cape.
eremobia Richards
Quartinia eremobia Richards, 1962, Revision. Study Masarid Wasps: 135 (key), 137, female, male –“Algeria: 80 Km. nord d'El Goléa” (holotype male, Paris); also from Libya: Tripolitania.
Distribution: Algeria; Libya.
eurydice (Richards), new combination
Quartinioides eurydice Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 183, female –“Cape Province: Bloukrans, near Calvinia” (London).
Distribution: South Africa: Northern Cape.
flava (Richards), new combination
Quartiniella flava Richards, 1962, Revision. Study Masarid Wasps: 208 (key), 210, female, male –“Cape Province: Nieuwoudtville” (holotype female, London).
Distribution: South Africa: Northern Cape.
funebris Kostylev
Quartinia funebris Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101, 103, female, male –“Imam-Baba (Transkaspien)” (St. Petersburg).
Distribution: Turkmenistan.
galataea (Richards), new combination
Quartinioides galataea Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 188, female –“Cape Province: Worcester” (London).
Distribution: South Africa: Western Cape.
goleana Richards
Quartinia goleana Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 137, female –“Algeria: 80 Km. nord d'El Goléa” (Paris).
Distribution: Algeria.
guichardi Richards
Quartinia guichardi Richards, 1969, J. Nat. Hist. 3: 80, female, male –“Canary Is., Gran Canaria, Maspalomas” (holotype male, London).
Distribution: Spain: Canary Islands: Gran Canaria.
haemorrhoa Gusenleitner
Quartinia haemorrhoa Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 137, 144, female –“Israel: Dead Sea, En Zeelim, 5 km N Massada, 31°23;pr/35°20;pr” (coll. M. Hauser).
Distribution: Israel.
halicticeps Giordani Soika
Quartinia halicticeps Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 1939: 9 (key), 12, figs. 2–4, female –“Egitto: Gabal Elba (Wadi Abrag)” (Venice, coll. Ministero de Agricoltura).
Distribution: Egypt.
hecuba (Richards), new combination
Quartinioides hecuba Richards, 1962, Revision. Study Masarid Wasps: 174 (key), 191, female, male –“Cape Province: Matjesfontein” (holotype male, London); also from Calvinia, and Kwazulu-Natal (“probably this species”).
Distribution: South Africa: Northern Cape, Western Cape, Kwazulu-Natal.
helena (Richards), new combination
Quartinioides helena Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 187, female –“Cape Province: Cape Town” (Washington).
Distribution: South Africa: Western Cape.
helichrysi (Richards), new combination
Quartinioides helichrysi Richards, 1962, Revision. Study Masarid Wasps: 175 (key), 198, female –“Basutoland: Mamalapi Mts.” (London).
Distribution: Lesotho.
hetaira (Richards), new combination
Quartinioides hetaira Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 200, female, male –“Cape Province: Cape Town” (holotype male, Washington).
Distribution: South Africa: Western Cape.
hypatia Richards
Quartinia hypatia Richards, 1962, Revision. Study Masarid Wasps: 130, 132 (key), 160, male –“Cape Province: Calvinia” (London).
Distribution: South Africa: Northern Cape.
indica Cameron
Juartinia[!] indica Cameron, 1904, Z. Hym. Dipt. 4: 89, male –“Deesa, Western India” (London).
Quartinia maculifrons Dover, 1925 (1924), J. Proc. Asiat. Soc. Bengal (n. ser.) 20(6): 290. Unavailable under Art. 11.5 of the International Code of Zoological Nomenclature.
Distribution: India: Gujarat (Deesa).
interrupta Turner
Quartinia interrupta Turner, 1939, Ann. Transvaal Mus. 20: 2, female –“Willowmore, Cape Province” (lectotype, Pretoria).
Distribution: Namibia; South Africa: Western Cape, Eastern Cape.
iphigenia (Richards), new combination
Quartinioides iphigenia Richards, 1962, Revision. Study Masarid Wasps: 173, 174, 175 (key), 192, male –“S.W. Africa: Aus” (London).
Distribution: Namibia.
jocasta Richards
Quartinia jocasta Richards, 1962, Revision. Study Masarid Wasps: 130 (key), 160, female –“Cape Province: Bloukrans, near Calvinia” (London); also from Calvinia.
Distribution: South Africa: Northern Cape.
laeta von Schulthess
Quartinia capensis var. læta von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 386 (key), 389, male, female –“Cape Province, Aus” (holotype male, London).
Distribution: Namibia.
latona (Richards), new combination
Quartinioides latona Richards, 1962, Revision. Study Masarid Wasps: 175, 177 (key), 207, female –“Cape Province: Prince Albert Road” (London).
Distribution: South Africa: Western Cape.
lesnei Benoist
Quartinia Lesnei Benoist, 1929, Bull. Soc. Sci. Nat. Maroc 9: 92, fig. 1, female –“Algérie: Tadmit” (Paris).
Distribution: Algeria.
libanica Richards
Quartinia libanica Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 83, male, female –“N. Lebanon: Becharré, 5000 ft.” (holotype male, Venice); also from elsewhere in Lebanon, and Cyprus.
Distribution: Cyprus; Lebanon.
maerens von Schulthess
Quartinia capensis var. mærens von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 386 (key), 389, female –“Cape Province, Worcester” (London); also from Matjesfontein.
Distribution: South Africa: Western Cape.
major Kohl
Quartinia major Kohl, 1898, Termesz. Füzetek 21: 363, pl. XV figs. 8, 21, female –“Oran … Auf Calendula” (Vienna).
?Quartinia kohli Dusmet, 1917, Mem. R. Soc. Españ. Hist. Nat. 8., Mem. 9: 383, male –“Imi-n-Tanut” (Madrid).
Distribution: Morocco; Algeria.
matabele Turner
Quartinia matabele Turner, 1939, Ann. Transvaal Mus. 20: 3, male, female –“Bulawayo, S. Rhodesia” (Pretoria).
Distribution: Zimbabwe.
media von Schulthess
Quartinia media von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 506, female –“Cape Province, Montagu” (London).
Distribution: South Africa: Western Cape.
medusa Richards
Quartinia medusa Richards, 1962, Revision. Study Masarid Wasps: 135 (key), 159, figs. 152–153, female, male –“S.W. Africa: Aus” (holotype male, London).
Distribution: Namibia.
metallescens von Schulthess
Quartinia metallescens von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 505, female –“Cape Province, Queenstown, 3500 feet” (London).
Distribution: South Africa: Eastern Cape; Lesotho.
metope (Richards), new combination
Quartinioides metope Richards, 1962, Revision. Study Masarid Wasps: 175, 178 (key), 207, female –“S.W. Africa: Aus” (London).
Distribution: Namibia.
minima von Schulthess
Quartinia minima von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 528, female –“Aus” (London).
Distribution: Namibia.
minuscula (Turner), new combination
Quartiniella minuscula Turner, 1939, Ann. Transvaal Mus. 20: 3, female –“Willowmore, Cape Province” (Pretoria).
Distribution: South Africa: Eastern Cape.
mochii Giordani Soika
Quartinia Mochii Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 1939: 9 (key), 10, fig. 1, male –“Egitto: Abu Rawash” (Venice, coll. Mochi).
Distribution: Egypt.
mongolica (Morawitz)
Celonites? mongolicus Morawitz, 1889, Horae Soc. Entomol. Ross. 23: 166, female, male –“Mongolia merid. Dshin-Tasy” (St. Petersburg).
Distribution: Mongolia.
multipicta (Richards), new combination
Quartinioides multipicta Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 202, female, male –“Cape Province: Worcester” (holotype male, London); also from elsewhere in Western Cape.
Distribution: South Africa: Western Cape.
nilotica Fischer
Quartinia nilotica Fischer, 1964, Ann. Naturhist. Mus. Wien 67: 611, fig. 1., female –“Khor Musa Pascha S. v. Wadi-Halfa” (Vienna).
Distribution: Egypt.
niveopicta von Schulthess
Quartinia niveopicta von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 327 (key), 329, female –“Cape Province, Matjestfontein” (London).
Distribution: South Africa: Western Cape, Eastern Cape.
nubiana Richards
Quartinia nubiana Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 138, female –“Egypt: Nubian Desert, Korosko to Abu Hamed” (London); also from elsewhere in Egypt.
Distribution: Tunisia; Libya; Egypt; Sudan; Saudi Arabia; United Arab Emirates.
ochraceopicta von Schulthess
Quartinia ochraceopicta von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 527, female –“Aus” (London).
Distribution: Namibia.
orientalis Gusenleitner
?“Quartinia haemorrhoa (n. sp.) frontalis n. ssp.” Blüthgen, 1961, Beitr. Naturk. Forsch. S.W.-Deutschl. 19(3): 281, male –“Gondahar.” Nomen nudum.
Quartinia orientalis Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 56, fig. 1b, male, female –“Türkei Ürgüp” (holotype male, Linz); also from Afghanistan.
Distribution: Turkey; Afghanistan.
paradoxa Brauns
Quartinia paradoxa Brauns, 1905, Ann. Mus. Natl. Hung. 3: 233, male –“Terra Capensis: Willowmore” (Pretoria).
Distribution: South Africa: Eastern Cape.
parcepunctata Richards
Quartinia parcepunctata Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 167, female, male –“Cape Town” (holotype female, Washington).
Distribution: South Africa: Western Cape.
parvula Dusmet
Quartinia parvula Dusmet, 1909, Act. Mem. Primer Congr. Nat. Españ., Zaragoza, 1908: 183, male, female (Madrid).
Distribution: Portugal; Spain.
perone Richards
Quartinia perone Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 166 –“Cape Province: Worcester” (London).
Distribution: South Africa: Western Cape.
persephone Richards
Quartinia persephone Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 165, female –“Cape Province: Calvinia” (London).
Distribution: South Africa: Northern Cape.
philomela (Richards), new combination
Quartinioides philomela Richards, 1962, Revision. Study Masarid Wasps: 173, 174 (key), 193, female –“Cape Province: Nieuwoudtville” (London).
Distribution: South Africa: Northern Cape.
phoebe (Richards), new combination
Quartinioides phoebe Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 181, male –“Cape Province: Cape Town” (Washington).
Distribution: South Africa: Western Cape.
pluto Richards
Quartinia pluto Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 166, female –“Cape Province: Worcester” (London).
Distribution: South Africa: Western Cape.
poecila von Schulthess
Quartinia poecila von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 327, fig. 2, female, male –“Cape Province, Swakopmund” (lectotype male, London).
Distribution: Namibia.
popovi Richards
Quartinia popovi Gussakovskii, 1935, Trudy Tadzik. Sect. Acad. Sci. U.S.S.R. (Zool. Parasitol.) 5: 466, fig. 23. Nomen nudum.
Quartinia popovi Richards, 1962, Revision. Study Masarid Wasps: 130 (key), 144, female –“District of Kondara” (Moscow).
Distribution: Tajikistan.
propinqua von Schulthess
Quartinia propinqua von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 526, figs. 2–4, male, female –“Aus” (lectotype female, London).
Distribution: Namibia; South Africa: Western Cape.
proserpina Richards
Quartinia proserpina Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 161, female –“Cape Province: Nieuwoudtville” (London).
Distribution: South Africa: Northern Cape.
punctulata von Schulthess
Quartinia punctulata von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 327 (key), 329, female –“Cape Province, Matjesfontein” (London).
Quartinia punctulatum[!] Turner, 1939, Ann. Transvaal Mus. 20: 1.
Distribution: South Africa: Western Cape.
pusilla Kostylev
Quartinia puslilla[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101. Incorrect original spelling.
Quartinia pusilla Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 103 (key), 104, female –“Krasnovodsk” (St. Petersburg).
Distribution: Turkmenistan.
senecionis (Richards), new combination
Quartinioides senecionis Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 205, female, male –“Orange Free State: Thaba Nchu” (holotype male, London); also from Lesotho.
Distribution: Lesotho; South Africa: Free State.
separata Gusenleitner
Quartinia separata Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 137, 146, female, male –“Syria west, 50 km S Homs” (holotype female, BOL); also from elsewhere in Syria; Turkey.
Distribution: Syria; Turkey.
shestakovi Kostylev
Quartinia shestakovi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101, 102 (key), 105, fig. 8, female, male –“Tartugaj (Syr-Darjja-Geb.)” (St. Petersburg); also from elsewhere in Uzbekistan.
Quartinia schestakovi[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 105. Incorrect original spelling.
Distribution: Uzbekistan; Kyrgyzstan.
signata von Schulthess
?Quartinia variegata “Brauns” Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 401. Nomen nudum.
“Quartinia signata, Brauns, in litt.” von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 507, male, female –“Cape Province, Willowmore” (lectotype male, Zürich).
Distribution: South Africa: Eastern Cape.
signatifrons Turner
Quartinia signata var. signatifrons Turner, 1939, Ann. Transvaal Mus. 20: 2, male, female (Pretoria).
Distribution: South Africa: Western Cape, Eastern Cape.
soikai Richards
Quartinia soikai Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 84, female –“Turkey: Steppe hills 70 m. S. of Ankara” (Venice).
Quartinia soika[!]; Gusenleitner, 1985, Nachricht. Bayer. Entomol. 34(4): 110.
Distribution: Iran; Turkey; Greece.
striata (Richards), new combination
Quartiniella striata Richards, 1962, Revision. Study Masarid Wasps: 208 (key), 211, female –“Cape Province: Calvinia” (London).
Distribution: South Africa: Northern Cape.
syriaca nitens Gusenleitner
Quartinia syriaca nitens Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 57, male –“Iran, Fars, Monti steppici 5 km N di Persepolis” (Venice).
Distribution: Iran.
syriaca syriaca Richards
Quartinia syriaca Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 86, female, male –“Syria: near Damascus, Ibn Nefis” (holotype female, Venice); also from other localities in Syria, and Lebanon.
Distribution: Lebanon; Syria.
tarsata (Richards), new combination
Quartinioides tarsata Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 185, female, male –“Cape Province: Grahamstown” (holotype female, London).
Distribution: South Africa: Eastern Cape.
tenerifina Richards
Quartinia tenerifina Richards, 1969, J. Nat. Hist. 3: 80, fig. 2, female, male –“Canary Is., Tenerife, Los Christianos” (holotype male, London).
Distribution: Spain: Canary Islands: Tenerife.
thebaica du Buysson
Quartinia thebaica du Buysson, 1902, Bull. Soc. Entomol. France 1902: 141, female –“Égypte: Louqsor” (coll. Pic).
Quartinia thebica[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 102.
Quartinia ochracea Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 1939: 9 (key), 14, female –“Egitto: Wadi Digla” (Venice, coll. Mochi).
Quartinia thabaica[!]; Gusenleitner, 1977, Linz. Biol. Beitr. 9: 152.
Distribution: Morocco; Algeria; Libya; Egypt.
titania (Richards), new combination
Quartinioides titania Richards, 1962, Revision. Study Masarid Wasps: 174 (key), 193, female, male –“Cape Province: Cape Town” (holotype male, Washington).
Distribution: South Africa: Western Cape.
tricolorata Giordani Soika
Quartinia thebaica du Buysson; Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 39: 13. Misidentification.
Quartinia tricolorata Giordani Soika, 1954, Boll. Mus. Civ. Stor. Nat. Venezia 7: 22, female –“Egitto: Gebel Asfar” (lectotype Venice); also from Fayum.
Distribution: Egypt.
tripolitana sinaitica Richards
Quartinia tripolitana sinaitica Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 83, male, female –“United Arab Republic, Sinai: Kun Mitla” (Venice).
Distribution: Egypt: Sinai.
tripolitana tripolitana Richards
Quartinia tripolitana Richards, 1962, Revision. Study Masarid Wasps: 134 (key), 140, male, female –“Tripolitania: Tripoli” (holotype male, London); also from elsewhere in Libya.
Distribution: Libya; Egypt.
tuareg Giordani Soika
Quartinia tuareg Giordani Soika, 1954, Boll. Mus. Civ. Stor. Nat. Venezia 7: 21, male, female –“Guelta Idjef Melène” (lectotype female, Venice); also from elsewhere in Algeria.
Distribution: Algeria; Egypt.
turneri (von Schulthess)
Quartiniella turneri von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 528, fig. 5, male –“Aus” (London).
Distribution: Namibia.
uzbeka Kostylev
Quartinia uzbeka Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101, 103 (key), 106, male –“Aman-Katan, Umgesb. v. Samarkand, 1.500 m. üb. M.” (St. Petersburg).
Distribution: Uzbekistan.
vagepunctata von Schulthess
Quartinia vagepunctata von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 507, female –“Cape Province, Little Karroo, 38 m. E. of Ceres” (London).
Distribution: South Africa: Northern Cape, Western Cape.
waterstoni (von Schulthess)
Quartiniella waterstoni von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 510, figs. 4–5, female, male –“Cape Province, Little Karroo, 38 m. E. of Ceres” (holotype female, London).
Distribution: South Africa: Northern Cape, Western Cape, Kwazulu-Natal.
Genus Trimeria de Saussure
Trimeria de Saussure, 1854, Ét. Fam. Vesp. 3: pl. IV fig. 2; 1855, Ét. Fam. Vesp. 3: 81, genus. Type species: Erynnis americana de Saussure, 1853, by monotypy.
Erynnis de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xix, genus (2 species), non Erynnis Schrank, 1801 (Lepidoptera). Type species: Erynnis americana de Saussure, 1853, by subsequent designation of de Saussure, 1855, Ét. Fam. Vesp. 3: 92.
Eurynnis[!] Dalla Torre, 1894, Cat. Hym. 9: 4.
Microtrimeria Bequaert, 1928, Psyche 35: 191, genus: 191. Type species: Microtrimeria cockerelli Bequaert, 1928, by monotypy.
Willinkia Richards, 1962, Revision. Study Masarid Wasps: 119, subgenus of Trimeria de Saussure. Type species: “Trimeria buyssoni Brèthes, 1904 (as represented in coll. B. M.),” by original designation.
americana (de Saussure)
Masaris vespiformis Fabricius; Romand, 1851, Ann. Soc. Entomol. France (2) 9, Bull.: lii (female not male). Misidentification.
Erynnis americana de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xx, female –“d'Amérique” (Paris).
Distribution: Brazil: Goiás,* Distrito Federal,* São Paulo,* Paraná.*
atacama (Fritz), new combination
Microtrimeria atacama Fritz, 1968, Rev. Chil. Entomol. 6: 69, female, male –“Estación Cachiyuyo, provincia de Atacama, Chile” (holotype female, Santiago).
Distribution: Chile: Atacama.
bequaerti Willink
Trimeria bequaerti Willink, 1951, Rev. Soc. Cient. Argentina 15: 78, figs. 1–6, female –“Amaicha del Valle, provincia de Tucumán (2.000 ms, s.n.m.)” (Tucumán); also from Salta.
Distribution: Bolivia; Paraguay: Boquerón; Argentina: Salta, Tucumán.
cockerelli (Bequaert), new combination
Microtrimeria cockerelli Bequaert, 1928, Psyche 35: 191, 192, female –“PERU: Payta” (Cambridge).
Distribution: Peru: Piura.
howardi Bertoni
Trimeria howardi Bertoni, 1911, An. Mus. Nac. Buenos Aires (3) 22: 104, female, male –“Puerto Bertoni” (San Lorenzo).
Distribution: Brazil*: Minas Gerais, São Paulo, Rio Grande do Sul; Paraguay: Alto Paraná; Argentina: Tucumán, Santa Fé, Misiones.
joergenseni Schrottky
Trimeria joergenseni Schrottky, 1909, Rev. Mus. La Plata 16: 137, female –“Argentina (Mendoza)” (destroyed).
Distribution: Argentina: Mendoza, San Luis,* Santa Fé,* Rio Negro.
monrosi Willink
Trimeria monrosi Willink, 1959, Neotropica 5: 2, female –“Tucumán, Las Cejas” (Tucumán).
Distribution: Bolivia: Santa Cruz; Argentina: Tucumán, Santiago del Estero,* Santa Fé.
neotropica (Mocsáry)
Jugurtia neotropica Mocsáry, 1906, Ann. Hist.-Nat. Mus. Hung. 4: 197, male –“Paraguay: Asuncion” (Budapest).
Distribution: Brazil*: Paraná; Paraguay: Central; Argentina*: Misiones.
rhachiphora (Schletterer)
Odynerus rhachiphorus Schletterer, 1891, Entomol. Nachr. 17: 85, male –“Chile” (Vienna).
Trimeria buyssoni Brèthes, 1903, An. Mus. Nac. Buenos Aires (3) 9(2): 371, figs. 1–2, 372, fig. 3, female –“Argentina: Catamarca” (Buenos Aires); also described from Córdoba? and Paraguay.
Trimeria rachiphorus[!]; Willink and Roig Alsina, 1993, J. Hym. Res. 2: 303.
Distribution: Paraguay: Boquerón, Itapuá; Uruguay*: Montevideo; Argentina: Jujuy, Salta, Tucumán, Catamarca, Santiago del Estero, Formosa, Santa Fé, Neuquén.
Acknowledgments
I thank Mike Engel, Fred Gess, Josef Gusenleitner, Norm Johnson, Lynn Kimsey, and Rob Longair for critical comments, and Vladimir Ovtsharenko for help with Russian localities.
REFERENCES
Appendices
INDEX OF MASARINAE
Genera, species, and subspecies are indexed, including emendations, misspellings, and lapsi calami, etc. The generic name given in parentheses following a species-group entry is the name used in the original combination. The name of the genus that now includes the species-group taxon is then listed below. If the species-group taxon is now considered a synonym, the valid name is listed following the current genus name. Similarly, for emendations, misspellings, and lapsi, the correct spelling follows. If the species-group taxon is considered a subspecies, this is indicated by the abbreviation “ssp.” following the name of the nominate species.
abbreviata (Vespa)
Celonites abbreviatus, 8
abbreviatus tauricus (Celonites)
Celonites abbreviatus ssp., 8
abbreviatus var. engadinensis (Celonites)
Celonites abbreviatus, 8
abbreviatus var. hungaricus (Celonites)
Celonites abbreviatus, 8
afer (Celonites)
Celonites 9
affinis (Quartinia)
Quartinia 22
albifrons (Pseudomasaris)
Pseudomasaris maculifrons 21
albopicta (Quartinioides)
Quartinia 22
alcestis (Quartinia)
Quartinia 22
alfkeni (Masaris)
Jugurtia 17
algerica (Masariella?)
Jugurtia 17
Alpha
Paragia 6
americana (Erynnis)
Trimeria 29
Ammoparagia
Metaparagia 5
andrei (Celonites)
Celonites 9
andromeda (Quartinioides)
Quartinia 22
angulata (Riekia)
Metaparagia 5
antennata (Quartinia)
Quartinia 23
antigone (Quartinioides)
Quartinia 23
apiformis (Masaris)
Celonites abbreviatus 8
araucana (Gayella)
Gayella 4
araxana (Quartinia)
Quartinia 23
arsinoe (Quartinioides)
Quartinia 23
artemis (Quartinia)
Quartinia 23
atacama (Microtrimeria)
Trimeria 30
atra (Quartinia)
Quartinia 23
atrata (Quartinia)
Quartinia 23
aucta (Liris)
Ceramius auctus 14
australis (Paragia)
Paragia 6
australis borealis (Paragia)
Paragia australis ssp. 6
bariscapus[!] (Pseudomasaris)
Pseudomasaris basirufus 21
bariscipus (Pseudomasaris)
Pseudomasaris basirufus 21
basuto (Quartinioides)
Quartinia 23
beaumonti (Paraceramius)
Ceramius 14
bequaerti (Trimeria)
Trimeria 30
bergenwahliae (Celonites)
Celonites 9
Beta
Paragia 6
beyeri (Ceramius)
Ceramius 14
bicolor (Paragia)
Paragia 6
bicolor (Philanthus)
Ceramius 14
bidens (Paragia)
Paragia odyneroides 7
bischoffi (Ceramius)
Ceramius 14
biskrensis (Jugurtia)
Jugurtia 17
borreriae (Rolandia)
Metaparagia 5
brasiliensis (Paramasaris)
Paramasaris 5
braunsi (Ceramiellus)
Jugurtia 17
braunsi (Ceramius)
Ceramius 14
braunsiella (Masaris)
Jugurtia 17
brevitarsis (Ceramius)
Ceramius 14
breyeri (Quartinia)
Quartinia 23
bureschi (Ceramius)
Ceramius 14
bureschi lycaonius (Ceramius)
Ceramius bureschi 14
buyssoni (Trimeria)
Trimeria rhachiphora 30
caffer (Ceramius)
Ceramius 14
calcarata (Jugurtia)
Jugurtia 17
calida (Paragia)
Paragia 6
canariensis (Quartinia)
Quartinia 23
candida (Quartinia)
Quartinia 23
capensis (Celonites)
Celonites 9
capensis (Ceramius)
Ceramius 14
capensis (Quartinia)
Quartinia 23
capensis var. laeta (Quartinia)
Quartinia laeta 25
capensis var. mærens (Quartinia)
Quartinia maerens 26
capicola (Ceramius)
Ceramius 14
carli (Masaris)
Masaris 20
caucasicus (Ceramius)
Ceramius 14
cazieri (Pseudomasaris)
Pseudomasaris 21
Celonites 8
Celonitinae
Masarinae 4
Ceramias[!]
Ceramius 13
Ceramiellus
Jugurtia 17
Ceramioides
Ceramius 13
Ceramiopsis 13
Ceramius 13
Ceramus[!]
Ceramius 13
cerceriformis (Ceramius)
Ceramius 14
cerceroides (Gayella)
Gayella patagonica 4
ceres (Masarina)
Masarina 19
ceres (Quartinioides)
Quartinia 23
Chelonites[!]
Celonites 8
chlorotica (Jugurtia)
Quartinia 23
cincta (Quartinia)
Quartinia 24
clarus (Celonites)
Celonites 9
clypeatus (Celonites)
Celonites 9
clypeatus (Ceramius)
Ceramius 15
cockerelli (Microtrimeria)
Trimeria 30
codoni (Jugurtia)
Jugurtia 17
Coelonites[!]
Celonites 8
communis (Masarina)
Masarina familiaris 19
concinna (Paragia)
Paragia venusta 8
confluens (Paragia)
Metaparagia 5
confusa (Jugurtia)
Jugurtia 17
coquilletti (Pseudomasaris)
Pseudomasaris 21
Cotyledon
Pseudomasaris 21
crenulatus (Celonites)
Celonites 9
cressida (Quartinioides)
Quartinia 24
cretica (Quartinia)
Quartinia 24
cupreus (Paramasaris)
Paramasaris 5
Cygnaea
Paragia 6
cyllene (Quartinioides)
Quartinia 24
cynara (Quartinioides)
Quartinia 24
cyprius (Celonites)
Celonites 9
cyprius smyrnensis (Celonites)
Celonites cyprius ssp. 9
damarinus (Ceramius)
Ceramius 15
davidi (Celonites)
Celonites 9
deceptor (Paragia)
Paragia 6
deceptrix (Paragia)
Paragia deceptor 6
decipiens (Paragia)
Paragia 7
decipiens aliciae (Paragia)
Paragia decipiens ssp. 6
diana (Quartinioides)
Quartinia 24
dilecta (Quartinia)
Quartinia 24
discrepans (Masaris)
Jugurtia 17
discretus (Celonites)
Celonites 9
dispar (Celonites)
Jugurtia 18
doddi (Metaparagia)
Metaparagia 6
dryope (Quartinioides)
Quartinia 24
dubia (Chrysis)
Celonites abbreviatus 8
duplicata (Jugurtia)
Jugurtia 18
edwardsii (Masaris)
Pseudomasaris 21
elissa (Quartinioides)
Quartinia 24
eremobia (Quartinia)
Quartinia 24
Erynnis
Trimeria 29
escalerae (Jugurtia)
Jugurtia 18
Eucelonites
Celonites 8
Euceramius
Ceramius 13
eumenoides (Gayella)
Gayella 4
eurycara (Jugurtia)
Jugurtia 18
eurydice (Quartinioides)
Quartinia 24
Eurynnis[!]
Trimeria 29
excellens (Paragia)
Paragia sobrina 8
familiaris (Masarina)
Masarina 19
fischeri (Celonites)
Celonites 9
flava (Quartiniella)
Quartinia 24
flavolineatus (Zethoides)
Paramasaris fuscipennis 5
fonscolombei (Ceramius)
Ceramius 15
fonscolombei oraniensis (Ceramius)
Ceramius fonscolombei ssp. 15
foveolatus (Celonites)
Celonites 9
foveolatus nigrior (Celonites)
Celonites foveolatus ssp. 9
fumipennis (Ceramius)
Ceramius linearis 15
funebris (Quartinia)
Quartinia 24
fuscipennis (Paramasaris)
Paramasaris 5
galataea (Quartinioides)
Quartinia 24
gariepensis (Celonites)
Celonites 9
Gayella 4
Gayellini 4
Gazella[!]
Gayella 4
generosa (Paragia)
Paragia 7
gessi (Ceramius)
Ceramius 15
gestroi (Ceramiopsis)
Ceramiopsis 13
Gnatho
Ceramius 13
goleana (Quartinia)
Quartinia 24
guichardi (Celonites)
Celonites 10
guichardi (Quartinia)
Quartinia 25
gussakovskii (Masaris)
Masaris 20
haemorrhoa (Quartinia)
Quartinia 25
haemorrhoa frontalis (Quartinia)
?Quartinia orientalis 27
halicticeps (Quartinia)
Quartinia 25
hamanni (Celonites)
Celonites 10
hammani[!] (Celonites)
Celonites hamanni 10
hecuba (Quartinioides)
Quartinia 25
helena (Quartinioides)
Quartinia 25
helichrysi (Quartinioides)
Quartinia 25
hellenicus (Celonites)
Celonites 10
hessei (Ceramius)
Ceramius nigripennis 16
hetaira (Quartinioides)
Quartinia 25
hirsuta (Paragia)
Paragia 7
hispanicus (Ceramius)
Ceramius 15
hoggarica (Jugurtia)
Jugurtia jemenensis 18
Holopticus
Pseudomasaris 21
houstoni (Rolandia)
Metaparagia 6
howardi (Trimeria)
Trimeria 30
hua (Ammoparagia)
Metaparagia 6
humeralis (Celonites)
Celonites 10
hyalinipennis (Masarina)
Masarina 19
hylaeiformis (Ceramius)
Masaris vespiformis 20
hypatia (Quartinia)
Quartinia 25
hystrix (Celonites)
Celonites 10
indica (Quartinia)
Quartinia 25
interrupta (Quartinia)
Quartinia 25
iphigenia (Quartinioides)
Quartinia 25
irana (Jugurtia)
Jugurtia 18
jacoti (Ceramius)
Ceramius 15
jemenensis (Jugurtia)
Jugurtia 18
jemenesis[!] (Jugurtia)
Jugurtia jemenensis 18
jocasta (Quartinia)
Quartinia 25
joergenseni (Trimeria)
Trimeria 30
joussaumei[!] (Celonites)
Celonites jousseaumei 10
jousseaumei (Celonites)
Celonites 10
jousseaumei asrensis (Celonites)
Celonites jousseaumei jousseaumei 10
jousseaumei senegalensis (Celonites)
Celonites jousseaumei ssp. 10
Juartinia[!]
Quartinia 22
Juguartia[!]
Jugurtia 17
Jugurthia[!]
Jugurtia 17
Jugurthia
Jugurtia 17
Jugurtia 17
Jujurtha[!]
Jugurtia 17
karooensis[!] (Ceramius)
Ceramius bicolor 14
karrooensis (Ceramius)
Ceramius bicolor 14
koeroegabensis (Jugurtia)
Jugurtia 18
kohli (Quartinia)
? Quartinia major 26
kostylevi (Celonites)
Celonites 10
kozlovi (Celonites)
Celonites 10
laetus (Celonites)
Celonites 10
latitarsis (Celonites)
Celonites 10
latona (Quartinioides)
Quartinia 25
lesnei (Quartinia)
Quartinia 25
libanica (Quartinia)
Quartinia 26
lichtensteinii (Gnatho)
Ceramius 15
linearis (Ceramius)
Ceramius 15
lobeliae (Celonites)
Celonites 10
longicornis (Saryara)
Masaris 20
longipilis (Celonites)
Celonites 11
luispenai (Gayella)
Gayella 4
lusitanicus (Ceramius)
Ceramius 15
lusitanicus var. luteo---clypeata (Ceramius)
Ceramius lusitanicus 15
macneilli (Pseudomasaris)
Pseudomasaris 21
macrocephalus (Ceramius)
Ceramius lichtensteinii 15
macswaini (Pseudomasaris)
Pseudomasaris 21
maculata (Paragia)
Metaparagia 6
maculifrons (Masaris)
Pseudomasaris 21
maculifrons (Quartinia)
Quartinia indica 25
magdalena (Paragia)
Paragia 7
major (Quartinia)
Quartinia 26
marginalis (Masaris)
Pseudomasaris 21
maroccanus (Paraceramius)
Ceramius 15
maroccanus rubripes (Ceramius)
Ceramius rubripes 16
Masariella
Jugurtia 17
Masarina 8 19
Masarini 5
Masaris 20
Massaris
Masaris 20
matabele (Quartinia)
Quartinia 26
mayeti (Celonites)
Celonites 11
Mazaridinae
Masarinae 4
Mazaris
Masaris 20
media (Quartinia)
Quartinia 26
medusa (Quartinia)
Quartinia 26
metallescens (Quartinia)
Quartinia 26
metanotalis (Ceramius)
Ceramius 16
Metaparagia 5
Metazethoides
Paramasaris 4
metope (Quartinioides)
Quartinia 26
michaelseni (Celonites)
Celonites 11
micheneri (Ceramius)
Ceramius 16
micheneri (Pseudomasaris)
Pseudomasaris 22
Microtrimeria
Trimeria 29
mimetica (Paragia)
Paragia 7
minima (Quartinia)
Quartinia 26
minuscula (Quartiniella)
Quartinia 26
mixta (Masarina)
Masarina 19
mixtoides (Masarina)
Masarina 19
mochii (Quartinia)
Quartinia 26
modestus (Celonites)
Celonites 11
modestus biinterruptus (Celonites)
Celonites modestus ssp. 11
modestus bisinterruptus[!] (Celonites)
Celonites modestus biinterruptus 11
mongolica (?Celonites)
Quartinia 26
monocesta (Paragia)
Paragia 7
monrosi (Trimeria)
Trimeria 30
montanus (Celonites)
Celonites 11
montanus (Ceramius)
Ceramius 16
morosa (Paragia)
Paragia 7
multipicta (Quartinioides)
Quartinia 26
mutilloides (Gayella)
Gayella 4
mutilloides nigerrima (Gayella)
Gayella mutilloides 4
nadigorum (Jugurtia)
Jugurtia 18
namaqua (Masarina)
Masarina 19
namibiensis (Priscomasaris)
Priscomasaris 5
nasuta (Paragia)
Paragia 7
neotropica (Jugurtia)
Trimeria 30
nigripennis (Ceramius)
Ceramius 16
nilotica (Quartinia)
Quartinia 26
niveopicta (Quartinia)
Quartinia 26
nocatunga (Riekia)
Metaparagia 6
nubiana (Quartinia)
Quartinia 26
numida[!] (Jugurtia)
Jugurtia oraniensis 18
numidica (Jugurtia)
Jugurtia oraniensis 18
nursei (Celonites)
Celonites 11
occidentalis (Masaris)
Pseudomasaris 22
ochracea (Quartinia)
Quartinia thebaica 28
ochraceopicta (Quartinia)
Quartinia 27
octoannulata (Saryara)
Celonites octoannulatus 11
octoannulatus hissaricus (Celonites)
Celonites octoannulatus ssp. 11
odyneroides (Gayella)
Gayella mutilloides 4
odyneroides (Paragia)
Paragia 7
oligomera (Paragia)
Paragia 7
oraniensis (Celonites)
Jugurtia 18
orientalis (Quartinia)
Quartinia 27
osseus (Celonites)
Celonites 11
palaestinensis (Paraceramius)
Ceramius 16
palestinensis[!] (Ceramius)
Ceramius palaestinensis 16
Paracelonites
Celonites 8
Paraceramius
Ceramius 13
paradoxa (Quartinia)
Quartinia 27
Paragayella
Paramasaris 5
Paragia 6
Paragiella
Paragia 6
Paragiina 5
paraguayensis (Ceramiopsis)
Ceramiopsis 13
Paramasaris 4
parcepunctata (Quartinia)
Quartinia 27
parvula (Masarina)
Masarina 19
parvula (Quartinia)
Quartinia 27
patagonica (Gayella)
Gayella 4
peliostomi (Celonites)
Celonites 11
peliostomi (Masarina)
Masarina 19
peringueyi (Ceramius)
Ceramius 16
perkinsi (Paragia)
Paragia 7
perone (Quartinia)
Quartinia 27
persephone (Quartinia)
Quartinia 27
persicus (Celonites)
Celonites 11
phaceliae (Pseudomasaris)
Pseudomasaris 22
philomela (Quartinioides)
Quartinia 27
phleomis]![ (Celonites)
Celonites phlomis 11
phlomis (Celonites)
Celonites 11
phoebe (Quartinioides)
Quartinia 27
pictifrons (Paragia)
Metaparagia 6
pictus (Celonites)
Celonites 11
pictus rufiventris (Celonites)
Celonites pictus ssp. 12
Plesiozethus
Paramasaris 4
pluto (Quartinia)
Quartinia 27
poecila (Quartinia)
Quartinia 27
popovi (Quartinia)
Quartinia 27
praedator (Paragia)
Paragia odyneroides 7
Priscomasaris 5
Priscomasarina 5
promontorii (Celonites)
Celonites 12
propinqua (Quartinia)
Quartinia 27
propodealis (Paragia)
Paragia 7
proserpina (Quartinia)
Quartinia 27
Pseudomasaris 21
punctulata (Quartinia)
Quartinia 27
purcelli (Celonites)
Celonites 12
purcelli var. flavomaculatus (Celonites)
Celonites purcelli 12
pusilla (Quartinia)
Quartinia 28
puslilla[!] (Quartinia)
Quartinia pusilla 28
Quartinia 22
Quartiniella
Quartinia 22
Quartinioides
Quartinia 22
rachiphorus[!] (Trimeria)
Trimeria rhachiphora 30
reedi (Gayella)
Gayella 4
rex (Ceramius)
Ceramius 16
rhachiphorus (Odynerus)
Trimeria rhachiphora 30
richardsi (Ceramius)
Ceramius 16
richardsi (Paragayella)
Paramasaris 5
Riekia
Metaparagia 5
rohweri (Pseudomasaris)
Pseudomasaris maculifrons 21
Rolandia
Metaparagia 5
romandi (Erynnis)
Masaris vespiformis 20
rothschildi (Celonites)
Celonites 12
rotschildi[!] (Celonites)
Celonites rothschildi 12
rudesculptus (Celonites)
Celonites 12
rufomaculatus (Ceramius)
Ceramius lichtensteinii 15
rugiceps (Celonites)
Celonites 12
Saryara
Masaris 20
Saryarinae
Masarinae 4
saussurei (Masaris)
Jugurtia 18
Masaris carli 20
saussurei[!] (Paragia)
Paragia tricolor 8
saussurii (Paragia)
Paragia tricolor 8
savignyi (Celonites)
?Celonites fischeri 9
schulthessi (Ceramius)
Ceramius cerceriformis 15
schulthessi (Paragia)
Paragia 7
scutellimacula (Quartinia)
Quartinia capensis 23
semenovi (Celonites)
Celonites 12
senecionis (Quartinioides)
Quartinia 28
separata (Quartinia)
Quartinia 28
shestakovi (Quartinia)
Quartinia 28
sicheliana (Gayella)
Gayella eumenoides 4
signata (Quartinia)
Quartinia 28
signata var. signatifrons (Quartinia)
Quartinia signatifrons 28
simpsoni (Jugurtia)
Jugurtia 18
smirnovi (Masaris)
Masaris carli 20
smithii (Paragia)
Paragia 7
sobrina (Paragia)
Paragia 8
socius (Ceramius)
Ceramius 16
soika[!] (Quartinia)
Quartinia soikai 28
soikai (Quartinia)
Quartinia 28
spinolae (Masaris)
Jugurtia 18
spinolai[!] (Jugurtia)
Jugurtia spinolae 19
spinosus (Celonites)
Celonites 12
spiricornis (Ceramius)
Ceramius auctus 14
spirolai[!] (Jugurtia)
Jugurtia spinolae 19
striata (Quartiniella)
Quartinia 28
strucki (Masarina)
Masarina 20
syriaca (Quartinia)
Quartinia 28
syriaca nitens (Quartinia)
Quartinia syriaca ssp. 28
tarsata (Quartinioides)
Quartinia 28
tenerifina (Quartinia)
Quartinia 28
testaceopicta (?Masariella)
Jugurtia simpsoni 18
texanus (Masaris)
Pseudomasaris 22
thabaica[!] (Quartinia)
Quartinia thebaica 29
thebaica (Quartinia)
Quartinia 28
thebica[!] (Quartinia)
Quartinia thebaica 28
tianshanicus (Masaris)
Masaris 20
tigrina (Jugurtia)
Jugurtia 19
titania (Quartinioides)
Quartinia 29
toriger (Ceramius)
Ceramius 16
Toryna
Pseudomasaris 21
tricolor (Paragia)
Paragia 8
tricolorata (Quartinia)
Quartinia 29
Trimeria 29
tripolitana (Quartinia)
Quartinia 29
tripolitana sinaitica (Quartinia)
Quartinia tripolitana ssp. 29
tristiculus (Celonites)
Celonites 12
tristiculus karatauicus (Celonites)
Celonites tristiculus ssp. 12
tuareg (Quartinia)
Quartinia 29
tuberculifer (Ceramius)
Ceramius 16
tumidiscutellatus (Celonites)
Celonites 12
turneri (?Masariella)
Jugurtia 19
turneri (Celonites)
Celonites 12
turneri (Quartiniella)
Quartinia 29
turneri eburnea (Masariella)
Jugurtia eburnea 18
tylecodoni (Masarina)
Masarina 20
uzbeka (Quartinia)
Quartinia 29
vagepunctata (Quartinia)
Quartinia 29
variegata (Quartinia)
? Quartinia signata 28
varipennis (Celonites)
Celonites 12
vechti (Ceramius)
Ceramius 17
venusta (Paragia)
Paragia 8
vespiformis (Ceramius)
Ceramius cerceriformis 15
vespiformis (Cimbex)
Celonites abbreviatus 8
vespiformis (Masaris)
Masaris 20
vespiformis (Paragia)
Paragia 8
vespiformis aegyptiacus (Masaris)
Masaris aegyptiacus 20
vespiformis arabicus (Masaris)
Masaris aegyptiacus ssp. 20
vespoides (Masaris)
Pseudomasaris 22
vespoides robertsoni (Pseudomasaris)
Pseudomasaris vespoides 22
wahlenbergiae (Celonites)
Celonites 13
walkeri (Paragia)
Paragia 8
waterstoni (Quartiniella)
Quartinia 29
wheeleri (Celonites)
Celonites 13
wheeleri (Pseudomasaris)
Pseudomasaris 22
wheeleri var. immaculatus (Celonites)
Celonites wheeleri 13
Willinkia
Trimeria 29
yemenensis (Celonites)
Celonites 13
yemenensis ethiopicus (Celonites)
Celonites yemenensis ssp. 13
yemensis[!] (Celonites)
Celonites 13
zarudnyi (Jugurtia)
Jugurtia 19
zavattarii (Celonites)
Celonites 13
Zethoides
Paramasaris 4
zonalis (Masaris)
Pseudomasaris 22
zonalis albopictus (Pseudomasaris)
Pseudomasaris zonalis 22
zonalis basirufus (Pseudomasaris)
Pseudomasaris basirufus 21
zonalis neomexicanus (Pseudomasaris)
Pseudomasaris maculifrons 21